Harumi Sakai, Yukimasa Ito, Sergei V. Shedko, Sergei N. Safronov, Sergei V. Frolov, Igor A. Chereshnev, Sang-Rin Jeon, Akira Goto: Phylogenetic and Taxonomic Relationships of Northern Far Eastern Phoxinin Minnows, Phoxinus and Rhynchocypris (Pisces, Cyprinidae), as Inferred from Allozyme and Mitochondrial 16S rRNA Sequence Analyses. In: Zoological Science. Band23, Nr.4, 2006, S.323–331 (englisch, biosoil.ru [PDF]).
