Matsuura, Keiichi (1 January 2015). "A new pufferfish of the genus Torquigener that builds "mystery circles" on sandy bottoms in the Ryukyu Islands, Japan (Actinopterygii: Tetraodontiformes: Tetraodontidae)". Ichthyological Research. 62 (2): 207–212. Bibcode:2015IchtR..62..207M. doi:10.1007/s10228-014-0428-5. ISSN1616-3915. S2CID254164102.
Matsuura, Keiichi (1 January 2015). "A new pufferfish of the genus Torquigener that builds "mystery circles" on sandy bottoms in the Ryukyu Islands, Japan (Actinopterygii: Tetraodontiformes: Tetraodontidae)". Ichthyological Research. 62 (2): 207–212. Bibcode:2015IchtR..62..207M. doi:10.1007/s10228-014-0428-5. ISSN1616-3915. S2CID254164102.
Matsuura, Keiichi (1 January 2015). "A new pufferfish of the genus Torquigener that builds "mystery circles" on sandy bottoms in the Ryukyu Islands, Japan (Actinopterygii: Tetraodontiformes: Tetraodontidae)". Ichthyological Research. 62 (2): 207–212. Bibcode:2015IchtR..62..207M. doi:10.1007/s10228-014-0428-5. ISSN1616-3915. S2CID254164102.
Matsuura, Keiichi (1 January 2015). "A new pufferfish of the genus Torquigener that builds "mystery circles" on sandy bottoms in the Ryukyu Islands, Japan (Actinopterygii: Tetraodontiformes: Tetraodontidae)". Ichthyological Research. 62 (2): 207–212. Bibcode:2015IchtR..62..207M. doi:10.1007/s10228-014-0428-5. ISSN1616-3915. S2CID254164102.