Rakotonasolo, Franck; Davis, Aaron (2006). "Six Species of Madagascan Genipa Transferred to Hyperacanthus (Rubiaceae-Gardenieae) and New Data on General Morphology, Placentation and Ovary Structure in Hyperacanthus". Taxon. 55 (2): 387–396. doi:10.2307/25065586. JSTOR25065586.
Rakotonasolo, Franck; Davis, Aaron (2006). "Six Species of Madagascan Genipa Transferred to Hyperacanthus (Rubiaceae-Gardenieae) and New Data on General Morphology, Placentation and Ovary Structure in Hyperacanthus". Taxon. 55 (2): 387–396. doi:10.2307/25065586. JSTOR25065586.