Le M, Raxworthy CJ, McCord WP, Mertz L (August 2006). "A molecular phylogeny of tortoises (Testudines: Testudinidae) based on mitochondrial and nuclear genes". Molecular Phylogenetics and Evolution. 40 (2): 517–31. doi:10.1016/j.ympev.2006.03.003. PMID16678445.
Pérez-García A, Vlachos E, Arribas A (March 2017). "The last giant continental tortoise of Europe: A survivor in the Spanish Pleistocene site of Fonelas P-1". Palaeogeography, Palaeoclimatology, Palaeoecology. 470: 30–39. Bibcode:2017PPP...470...30P. doi:10.1016/j.palaeo.2017.01.011. hdl:10261/277114.
Pérez-García A, Vlachos E, Arribas A (March 2017). "The last giant continental tortoise of Europe: A survivor in the Spanish Pleistocene site of Fonelas P-1". Palaeogeography, Palaeoclimatology, Palaeoecology. 470: 30–39. Bibcode:2017PPP...470...30P. doi:10.1016/j.palaeo.2017.01.011. hdl:10261/277114.
Pérez-García A, Vlachos E, Arribas A (March 2017). "The last giant continental tortoise of Europe: A survivor in the Spanish Pleistocene site of Fonelas P-1". Palaeogeography, Palaeoclimatology, Palaeoecology. 470: 30–39. Bibcode:2017PPP...470...30P. doi:10.1016/j.palaeo.2017.01.011. hdl:10261/277114.
Le M, Raxworthy CJ, McCord WP, Mertz L (August 2006). "A molecular phylogeny of tortoises (Testudines: Testudinidae) based on mitochondrial and nuclear genes". Molecular Phylogenetics and Evolution. 40 (2): 517–31. doi:10.1016/j.ympev.2006.03.003. PMID16678445.