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Embley, T. Martin; William Martin (2006). "Eukaryotic evolution, changes and challenges". Nature440 (7084): 623–630. PMID16572163. doi:10.1038/nature04546.
Bell, A.C.; G. Felsenfeld (2000). "Methylation of a CTCF-dependent boundar control imprinted expression of the Igf2 gene". Nature405 (6785): 482–485. PMID10839546. doi:10.1038/35013100.