U. Fritz, A. K. Hundsdörfer, P. Široký, M. Auer, H. Kami, J. Lehmann, L. F. Mazanaeva, O. Türkozan, M. Wink: Phenotypic plasticity leads to incongruence between morphology-based taxonomy and genetic differentiation in western Palaearctic tortoises (Testudo graeca complex; Testudines, Testudinidae). In: Amphibia-Reptilia. 28 (1), 2007, S. 7–121 (ZusammenfassungArchiválva2007. október 2-i dátummal a Wayback Machine-ben)
Uwe Fritz, D. James Harris, Soumia Fahd, Rachid Rouag, Eva Graciá Martínez, Andrés Giménez Casalduero, Pavel Široký, Mohsen Kalboussi, Tarek B. Jdeidi, Anna K. Hundsdörfer (2009): Mitochondrial phylogeography of Testudo graeca in the Western Mediterranean: Old complex divergence in North Africa and recent arrival in Europe.- Amphibia-Reptilia 30 (2009): 63-80 Zusammenfassung[halott link]
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James F. Parham, Oğuz Türkozan, Bryan L. Stuart, Marine Arakelyan, Soheila Shafel, J. Robert Macey, Yehudah L. Werner, Theodore J. Papenfuss: Genetic Evidence for Premature Taxonomic Inflation in Middle Eastern Tortoises. In: Proceedings of the California Academy of Sciences. Fourth Series, Band 57, Nr. 3, 2006, ISSN0068-547X, S. 955–964
U. Fritz, A. K. Hundsdörfer, P. Široký, M. Auer, H. Kami, J. Lehmann, L. F. Mazanaeva, O. Türkozan, M. Wink: Phenotypic plasticity leads to incongruence between morphology-based taxonomy and genetic differentiation in western Palaearctic tortoises (Testudo graeca complex; Testudines, Testudinidae). In: Amphibia-Reptilia. 28 (1), 2007, S. 7–121 (ZusammenfassungArchiválva2007. október 2-i dátummal a Wayback Machine-ben)
Uwe Fritz, D. James Harris, Soumia Fahd, Rachid Rouag, Eva Graciá Martínez, Andrés Giménez Casalduero, Pavel Široký, Mohsen Kalboussi, Tarek B. Jdeidi, Anna K. Hundsdörfer (2009): Mitochondrial phylogeography of Testudo graeca in the Western Mediterranean: Old complex divergence in North Africa and recent arrival in Europe.- Amphibia-Reptilia 30 (2009): 63-80 Zusammenfassung[halott link]