Villalobos, Federico (2013년 6월 1일). “Tree squirrels: A key to understand the historic biogeography of Mesoamerica?”. 《Mammalian Biology - Zeitschrift für Säugetierkunde》 78 (4): 258–266. doi:10.1016/j.mambio.2013.02.003.
Pečnerová, Patrícia; Moravec, Jiří C.; Martínková, Natália (2015). “A Skull Might Lie: Modeling Ancestral Ranges and Diet from Genes and Shape of Tree Squirrels”. 《Systematic Biology》 64 (6): 1074. doi:10.1093/sysbio/syv054. PMID26254670.