Wilson Don E. & Reeder DeeAnn M. (red.) Gulo. w: Mammal Species of the World. A Taxonomic and Geographic Reference (Wyd. 3.) [on-line]. Johns Hopkins University Press, 2005. (ang.) [dostęp 2008-06-08]
doi.org
dx.doi.org
ChapronCh.G.ChapronCh. i inni, Recovery of large carnivores in Europe’s modern human-dominated landscapes, „Science”, 346 (6216), 2014, s. 1517–1519, DOI: 10.1126/science.1257553.i inni
doi.org
Joshua X. Samuels, Keila E. Bredehoeft i Steven C. Wallace. A new species of Gulo from the Early Pliocene Gray Fossil Site (Eastern United States); rethinking the evolution of wolverines. „PeerJ”. 6:e4648, 2018. DOI: 10.7717/peerj.4648. (ang.).
europa.eu
ec.europa.eu
Gulo gulo [online], IUCN 2007. European Mammal Assessment [dostęp 2008-06-09] [zarchiwizowane z adresu 2008-07-19](ang.).