Guayasamin J. M., Krynak T., Krynak K., Culebras J. and Hutter C. R. Phenotypic plasticity raises questions for taxonomically important traits: a remarkable new Andean rainfrog (Pristimantis) with the ability to change skin texture (англ.) // Zoological Journal of the Linnean Society. — 2005. — Vol. 173, no. 4. — P. 913—928. — doi:10.1111/zoj.12222.