Richard G. Hartnoll; Mark S. P. Baine; Yolima Grandas; Jennifer James & Helen Atkin. Population biology of the black land crab, Gecarcinus ruricola, in the San Andres archipelago, Western Caribbean. Journal of Crustacean Biology. 2006, 26 (3): 316–325. doi:10.1651/C-2640.1.
Why do we see Crabs in the Quill?(PDF). St Eustatius: National and Marine Parks and Botanical Gardens Newsletter. 2009: 5 [2019-02-20]. (原始内容(PDF)存档于2012-02-24).
Why do we see Crabs in the Quill?(PDF). St Eustatius: National and Marine Parks and Botanical Gardens Newsletter. 2009: 5 [2019-02-20]. (原始内容(PDF)存档于2012-02-24).