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Kimbel، W؛ Lockwood، C؛ Ward، C؛ Leakey، M؛ Rak، Y؛ Johanson، D (2006). "Was Australopithecus anamensis ancestral to A. Afarensis? A case of anagenesis in the hominin fossil record". Journal of Human Evolution. ج. 51 ع. 2: 134–52. DOI:10.1016/j.jhevol.2006.02.003. PMID:16630646.