Andreas R.Dunz & Ulrich K.Schliewen (2013). "Molecular phylogeny and revised classification of the haplotilapiine cichlid fishes formerly referred to as Tilapia". Molecular Phylogenetics and Evolution. ج. 68 ع. 1: 64–80. DOI:10.1016/j.ympev.2013.03.015. PMID:23542002.
Andreas R.Dunz & Ulrich K.Schliewen (2013). "Molecular phylogeny and revised classification of the haplotilapiine cichlid fishes formerly referred to as Tilapia". Molecular Phylogenetics and Evolution. ج. 68 ع. 1: 64–80. DOI:10.1016/j.ympev.2013.03.015. PMID:23542002.