Emerald B.S.؛ Roy J. K. (1998). "Organising activities of engrailed, hedgehog, wingless and decapentaplegic in the genital discs of Drosophila melanogaster". Development Genes and Evolution. ج. 208: 504–516. DOI:10.1007/s004270050209.
Tsuda M، Kamimura K، Archer M، Fox B، Olson S (1999). "The cell-surface proteoglican Dally regulates Wingless signaling in Drosophila". Nature. ج. 400 ع. 6741: 276–280. DOI:10.1038/22336. PMID:10421371.