Razafimandimbison SG، Lantz H، Mouly A، Bremer B (2009). "Evolutionary trends, major lineages, and new generic limits in the dioecious group of the tribe Vanguerieae (Rubiaceae): insights into the evolution of functional dioecy". Annals of the Missouri Botanical Garden. ج. 96 ع. 1: 161–181. DOI:10.3417/2006191. S2CID:86065468.
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Razafimandimbison SG، Lantz H، Mouly A، Bremer B (2009). "Evolutionary trends, major lineages, and new generic limits in the dioecious group of the tribe Vanguerieae (Rubiaceae): insights into the evolution of functional dioecy". Annals of the Missouri Botanical Garden. ج. 96 ع. 1: 161–181. DOI:10.3417/2006191. S2CID:86065468.
