van Dijk, Peter Paul; Iverson, John B.; Shaffer, H. Bradley; Bour, Roger; Rhodin, Anders G. J.. «Turtles of the World, 2011 Update: Annotated Checklist of Taxonomy, Synonymy, Distribution, and Conservation Status». A: Conservation Biology of Freshwater Turtles and Tortoises, 2011, p. 000.197. DOI10.3854/crm.5.000.checklist.v4.2011. OCLC472656069.
Edwards, Danielle; Benavides, Edgar; Garrick, Ryan «The genetic legacy of Lonesome George survives: Giant tortoises with Pinta Island ancestry identified in Galápagos». Biological Conservation, 157, 1-2013, pàg. 225–228. DOI: 10.1016/j.biocon.2012.10.014.
van Dijk, Peter Paul; Iverson, John B.; Shaffer, H. Bradley; Bour, Roger; Rhodin, Anders G. J.. «Turtles of the World, 2011 Update: Annotated Checklist of Taxonomy, Synonymy, Distribution, and Conservation Status». A: Conservation Biology of Freshwater Turtles and Tortoises, 2011, p. 000.197. DOI10.3854/crm.5.000.checklist.v4.2011. OCLC472656069.