Artiss T, Schultz TR, Polhemus DA, Simon C «Molecular phylogenetic analysis of the dragonfly genera Libellula, Ladona, and Plathemis (Odonata: Libellulidae) based on mitochondrial cytochrome oxidase I and 16S rRNA sequence data». Molecular Phylogenetics and Evolution, 18, 3, 2001, pàg. 348–61. DOI: 10.1006/mpev.2000.0867. PMID: 11277629.
Kambhampati, Srinivas; Charlton, Ralph E. «Phylogenetic relationship among Libellula, Ladona and Plathemis (Odonata: Libellulidae) based on DNA sequence of mitochondrial 16S rRNA gene». Systematic Entomology, 24, 1, 1999, pàg. 37–49. DOI: 10.1046/j.1365-3113.1999.00066.x.
Artiss T, Schultz TR, Polhemus DA, Simon C «Molecular phylogenetic analysis of the dragonfly genera Libellula, Ladona, and Plathemis (Odonata: Libellulidae) based on mitochondrial cytochrome oxidase I and 16S rRNA sequence data». Molecular Phylogenetics and Evolution, 18, 3, 2001, pàg. 348–61. DOI: 10.1006/mpev.2000.0867. PMID: 11277629.