Danmarks Fauna. Victor Hansen og Sv. G Larsson. Biller XIX. Almindelig del GEC Gads Forlag. København 1957.
McHugh, Joseph V.; Liebherr, James K. (2009). "Coleoptera". I Vincent H. Resh; Ring T. Cardé (red.). Encyclopedia of Insects (2nd udgave). Academic Press. s. 183–201. ISBN978-0-12-374144-8.
Hunt, T.; Bergsten, J.; Levkanicova, Z.; Papadopoulou, A.; John, O. S.; Wild, R.; Hammond, P. M.; Ahrens, D.; Balke, M.; Caterino, M. S.; Gómez-Zurita, J.; Ribera, I; Barraclough, T. G.; Bocakova, M.; Bocak, L; Vogler, A. P. (2007). "A Comprehensive Phylogeny of Beetles Reveals the Evolutionary Origins of a Superradiation". Science. 318 (5858): 1913-1916. Bibcode:2007Sci...318.1913H. doi:10.1126/science.1146954. ISSN0036-8075. PMID18096805. S2CID19392955.
Hunt, T.; Bergsten, J.; Levkanicova, Z.; Papadopoulou, A.; John, O. S.; Wild, R.; Hammond, P. M.; Ahrens, D.; Balke, M.; Caterino, M. S.; Gómez-Zurita, J.; Ribera, I; Barraclough, T. G.; Bocakova, M.; Bocak, L; Vogler, A. P. (2007). "A Comprehensive Phylogeny of Beetles Reveals the Evolutionary Origins of a Superradiation". Science. 318 (5858): 1913-1916. Bibcode:2007Sci...318.1913H. doi:10.1126/science.1146954. ISSN0036-8075. PMID18096805. S2CID19392955.
Hunt, T.; Bergsten, J.; Levkanicova, Z.; Papadopoulou, A.; John, O. S.; Wild, R.; Hammond, P. M.; Ahrens, D.; Balke, M.; Caterino, M. S.; Gómez-Zurita, J.; Ribera, I; Barraclough, T. G.; Bocakova, M.; Bocak, L; Vogler, A. P. (2007). "A Comprehensive Phylogeny of Beetles Reveals the Evolutionary Origins of a Superradiation". Science. 318 (5858): 1913-1916. Bibcode:2007Sci...318.1913H. doi:10.1126/science.1146954. ISSN0036-8075. PMID18096805. S2CID19392955.
Hunt, T.; Bergsten, J.; Levkanicova, Z.; Papadopoulou, A.; John, O. S.; Wild, R.; Hammond, P. M.; Ahrens, D.; Balke, M.; Caterino, M. S.; Gómez-Zurita, J.; Ribera, I; Barraclough, T. G.; Bocakova, M.; Bocak, L; Vogler, A. P. (2007). "A Comprehensive Phylogeny of Beetles Reveals the Evolutionary Origins of a Superradiation". Science. 318 (5858): 1913-1916. Bibcode:2007Sci...318.1913H. doi:10.1126/science.1146954. ISSN0036-8075. PMID18096805. S2CID19392955.
Hunt, T.; Bergsten, J.; Levkanicova, Z.; Papadopoulou, A.; John, O. S.; Wild, R.; Hammond, P. M.; Ahrens, D.; Balke, M.; Caterino, M. S.; Gómez-Zurita, J.; Ribera, I; Barraclough, T. G.; Bocakova, M.; Bocak, L; Vogler, A. P. (2007). "A Comprehensive Phylogeny of Beetles Reveals the Evolutionary Origins of a Superradiation". Science. 318 (5858): 1913-1916. Bibcode:2007Sci...318.1913H. doi:10.1126/science.1146954. ISSN0036-8075. PMID18096805. S2CID19392955.