T. D. Grant, D. R. Frost, J. P. Caldwell, R. Gagliardo, C. F. B. Haddad, P. J. R. Kok, B. D. Means, B. P. Noonan, W. Schargel & W. C. Wheeler: Phylogenetic systematics of dart-poison frogs and their relatives (Amphibia: Athesphatanura: Dendrobatidae). Bulletin of the American Museum of Natural History, 299, S. 173, 2006 PDF, online
T. D. Grant, D. R. Frost, J. P. Caldwell, R. Gagliardo, C. F. B. Haddad, P. J. R. Kok, B. D. Means, B. P. Noonan, W. Schargel & W. C. Wheeler: Phylogenetic systematics of dart-poison frogs and their relatives (Amphibia: Athesphatanura: Dendrobatidae). Bulletin of the American Museum of Natural History, 299, S. 111–114, 2006 PDF, online
T. D. Grant, D. R. Frost, J. P. Caldwell, R. Gagliardo, C. F. B. Haddad, P. J. R. Kok, B. D. Means, B. P. Noonan, W. Schargel & W. C. Wheeler: Phylogenetic systematics of dart-poison frogs and their relatives (Amphibia: Athesphatanura: Dendrobatidae). Bulletin of the American Museum of Natural History, 299, S. 178, 2006 PDF, online
research.amnh.org
Darrel R. Frost: Dendrobates. Amphibian Species of the World: an Online Reference. Electronic Database accessible at American Museum of Natural History, Version 5.5 vom 31. Januar 2011, New York 1998–2011
archive.org
Johann Georg Wagler: Natürliches System der Amphibien : mit vorangehender Classification der Säugethiere und Vögel : ein Beitrag zur vergleichenden Zoologie. München 1830, S. 202 (Erstbeschreibung) Faksimile bei Internet Archive