Argolo, L.A., López-Fernández, H., Batalha-Filho, H. & Mello Affonso, P.R.A. (2020): Unraveling the systematics and evolution of the ‘Geophagus’ brasiliensis (Cichliformes: Cichlidae) species complex. Molecular Phylogenetics and Evolution, 150: 106855. doi: 10.1016/j.ympev.2020.106855
Hernán López-Fernández, Kirk O. Winemiller & Rodney L. Honeycutt: Multilocus phylogeny and rapid radiations in Neotropical cichlid fishes (Perciformes: Cichlidae: Cichlinae). Molecular Phylogenetics and Evolution Volume 55, Issue 3, June 2010, Pages 1070-1086 doi:10.1016/j.ympev.2010.02.020