T. Grant, D.R. Frost, J.P. Caldwell, R. Gagliardo, C.F.B. Haddad, P.J.R. Kok, D.B. Means, B.P. Noonan, W.E Schargel & W.C. Wheeler (2006): Phylogenetic systematics of dart-poison frogs and their relatives (Amphibia: Athesphatanura: Dendrobatidae). Bulletin of the American Museum of Natural History 299, S. 165 PDF online
