Thomas J. Simonsen, Rienk de Jong, Maria Heikkilä, Lauri Kaila (2012): Butterfly morphology in a molecular age. Does it still matter in butterfly systematics? Arthropod Structure & Development 41: 307-322. doi:10.1016/j.asd.2012.04.006
Jerome C. Regier, Charles Mitter u. a.: A Large-Scale, Higher-Level, Molecular Phylogenetic Study of the Insect Order Lepidoptera (Moths and Butterflies). In: PLoS ONE. 8, 2013, S. e58568, doi:10.1371/journal.pone.0058568.
