Valiela, Ivan (1974). «Composition, food webs, and population limitation in dung arthropod communities during invasion and succession». American Midland Naturalist92: 370-385 [380]. doi:10.2307/2424302.
Finn, J.A.; Gittings, T. (2003). «A review of competition in north temperate dung beetle communities». Ecological Entomology28 (1): 1-13. doi:10.1046/j.1365-2311.2002.00487.x.