Pemadasa, M. A.; Greig-Smith, P.; Lovell, P. H. (July 1974). "A Quantitative Description of the Distribution of Annuals in the Dune System at Aberffraw, Anglesey". Journal of Ecology. 62 (2): 379–402. doi:10.2307/2258986. JSTOR2258986.
Pemadasa, M. A.; Greig-Smith, P.; Lovell, P. H. (July 1974). "A Quantitative Description of the Distribution of Annuals in the Dune System at Aberffraw, Anglesey". Journal of Ecology. 62 (2): 379–402. doi:10.2307/2258986. JSTOR2258986.