Sanborn, Colin Campbell. Descriptions and records of African bats. Zoological series. Vol. 20. Field Museum of Natural History. pp. 111–112. Retrieved October 9, 2017.
Lutz, H. L.; Patterson, B. D.; Peterhans, J. C. K.; Stanley, W. T.; Webala, P. W.; Gnoske, T. P.; Hackett, S.J.; Stanhope, M. J. (2016). "Diverse sampling of East African haemosporidians reveals chiropteran origin of malaria parasites in primates and rodents". Molecular Phylogenetics and Evolution. 99: 7–15. doi:10.1016/j.ympev.2016.03.004. PMID26975691.
Lutz, H. L.; Patterson, B. D.; Peterhans, J. C. K.; Stanley, W. T.; Webala, P. W.; Gnoske, T. P.; Hackett, S.J.; Stanhope, M. J. (2016). "Diverse sampling of East African haemosporidians reveals chiropteran origin of malaria parasites in primates and rodents". Molecular Phylogenetics and Evolution. 99: 7–15. doi:10.1016/j.ympev.2016.03.004. PMID26975691.