Gilles A; Lecointre G; Faure E; Chappaz R; Brun G. (1998). "Mitochondrial phylogeny of the European cyprinids: implications for their systematics, reticulate evolution, and colonization time (abstract)". Molecular Phylogenetics and Evolution. 10 (1): 132–143. doi:10.1006/mpev.1997.0480. PMID9751923.
Gilles A; Lecointre G; Faure E; Chappaz R; Brun G. (1998). "Mitochondrial phylogeny of the European cyprinids: implications for their systematics, reticulate evolution, and colonization time (abstract)". Molecular Phylogenetics and Evolution. 10 (1): 132–143. doi:10.1006/mpev.1997.0480. PMID9751923.