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Cole, C. J.; Townsend, C. R.; Reynolds, R. P.; MacCulloch, R. D.; Lathrop, A. (2013). "Amphibians and reptiles of Guyana, South America: Illustrated keys, annotated species accounts, and a biogeographic synopsis". Proceedings of the Biological Society of Washington. 125 (4): 317–578. doi:10.2988/0006-324X-125.4.317. S2CID86665287.
Cole, C. J.; Townsend, C. R.; Reynolds, R. P.; MacCulloch, R. D.; Lathrop, A. (2013). "Amphibians and reptiles of Guyana, South America: Illustrated keys, annotated species accounts, and a biogeographic synopsis". Proceedings of the Biological Society of Washington. 125 (4): 317–578. doi:10.2988/0006-324X-125.4.317. S2CID86665287.