Serrano-Rojas, Shirley J.; Whitworth, Andrew; Villacampa, Jaime; May, Rudolf Von; Gutiérrez, Roberto C.; Padial, José M.; Chaparro, Juan C. (31 December 2017). "A new species of poison-dart frog (Anura: Dendrobatidae) from Manu province, Amazon region of southeastern Peru, with notes on its natural history, bioacoustics, phylogenetics, and recommended conservation status". Zootaxa. 4221 (1): 71–94. doi:10.11646/zootaxa.4221.1.3. PMID28187674.
Serrano-Rojas, Shirley J.; Whitworth, Andrew; Villacampa, Jaime; May, Rudolf Von; Gutiérrez, Roberto C.; Padial, José M.; Chaparro, Juan C. (31 December 2017). "A new species of poison-dart frog (Anura: Dendrobatidae) from Manu province, Amazon region of southeastern Peru, with notes on its natural history, bioacoustics, phylogenetics, and recommended conservation status". Zootaxa. 4221 (1): 71–94. doi:10.11646/zootaxa.4221.1.3. PMID28187674.