Anatomy of pterosaurs (English Wikipedia)

Analysis of information sources in references of the Wikipedia article "Anatomy of pterosaurs" in English language version.

Last modified: 05:22:58, 14 April 2026 (UTC)

Ref.Un. Ref.Website

Global rank English rank

4134doi.org

2^nd place

3327nih.gov

5^th place

2622adsabs.harvard.edu

14^th place

9^th place

1816semanticscholar.org

15^th place

8^th place

33rero.ch

3,023^rd place

2,092^nd place

33hdl.handle.net

66^th place

34^th place

32worldcat.org

4^th place

32biodiversitylibrary.org

168^th place

136^th place

21oup.com

292^nd place

233^rd place

11nature.com

207^th place

347^th place

11sciencedirect.com

137^th place

146^th place

11researchgate.net

96^th place

89^th place

11bris.ac.uk

5,759^th place

4,110^th place

11bbc.com

26^th place

23^rd place

adsabs.harvard.edu (Global: 14^th place; English: 9^th place)

ui.adsabs.harvard.edu

Witmer LM, Chatterjee S, Franzosa J, Rowe T (2003). "Neuroanatomy of flying reptiles and implications for flight, posture and behaviour" (PDF). Nature. 425 (6961): 950–53. Bibcode:2003Natur.425..950W. doi:10.1038/nature02048. PMID 14586467. S2CID 4431861.
Etienne, J. L.; Smith, R. E.; Unwin, D. M.; Smyth, R. S. H.; Martill, D. M. (2024). "A 'giant' pterodactyloid pterosaur from the British Jurassic". Proceedings of the Geologists' Association. 135 (3): 335–348. Bibcode:2024PrGA..135..335E. doi:10.1016/j.pgeola.2024.05.002.
Jagielska, N.; Challands, T. J.; O'Sullivan, M.; Ross, D. A.; Fraser, N. C.; Wilkinson, M.; Brusatte, S. L. (2023). "New postcranial remains from the Lealt Shale Formation of the Isle of Skye, Scotland, showcase hidden pterosaur diversity in the Middle Jurassic". Scottish Journal of Geology. 59 (1–2): 001. Bibcode:2023ScJG...59....1J. doi:10.1144/sjg2023-001. hdl:20.500.11820/8bc004a4-ab80-4f9f-965d-f211f18e9876. S2CID 258232744.
Martin-Silverstone, Elizabeth; Witton, Mark P.; Arbour, Victoria M.; Currie, Philip J. (2016). "A small azhdarchoid pterosaur from the latest Cretaceous, the age of flying giants". Royal Society Open Science. 3 (8) 160333. Bibcode:2016RSOS....360333M. doi:10.1098/rsos.160333. PMC 5108964. PMID 27853614.
Hone, D.W.E. (2020). "A review of the taxonomy and palaeoecology of the Anurognathidae (Reptilia, Pterosauria)". Acta Geologica Sinica - English Edition. 94 (5): 1676–1692. Bibcode:2020AcGlS..94.1676H. doi:10.1111/1755-6724.14585.
Naish, D; Witton, MP; Martin-Silverstone, E (2021). "Powered flight in hatchling pterosaurs: evidence from wing form and bone strength". Scientific Reports. 11 (1) 13130. Bibcode:2021NatSR..1113130N. doi:10.1038/s41598-021-92499-z. PMC 8298463. PMID 34294737.
Witton, M.P.; Habib, M.B. (2010). "On the Size and Flight Diversity of Giant Pterosaurs, the Use of Birds as Pterosaur Analogues and Comments on Pterosaur Flightlessness". PLOS ONE. 5 (11) e13982. Bibcode:2010PLoSO...513982W. doi:10.1371/journal.pone.0013982. PMC 2981443. PMID 21085624.
Andres, B.; Langston, W. Jr. (2021). "Morphology and taxonomy of Quetzalcoatlus Lawson 1975 (Pterodactyloidea: Azhdarchoidea)". Journal of Vertebrate Paleontology. 41 (sup1): 46–202. Bibcode:2021JVPal..41S..46A. doi:10.1080/02724634.2021.1907587. ISSN 0272-4634. S2CID 245125409.
Frey E, Martill DM (1998). "Soft tissue preservation in a specimen of Pterodactylus kochi (Wagner) from the Upper Jurassic of Germany". Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen. 210 (3): 421–41. Bibcode:1998NJGPA.210..421F. doi:10.1127/njgpa/210/1998/421.
Andres, B.; Ji Q. (2008). "A new pterosaur from the Liaoning Province of China, the phylogeny of the Pterodactyloidea, and convergence in their cervical vertebrae". Palaeontology. 51 (2): 453–469. Bibcode:2008Palgy..51..453A. doi:10.1111/j.1475-4983.2008.00761.x.
Andres, B.; Clark, J. M.; Xing, X. (2010). "A new rhamphorhynchid pterosaur from the Upper Jurassic of Xinjiang, China, and the phylogenetic relationships of basal pterosaurs" (PDF). Journal of Vertebrate Paleontology. 30 (1): 163–187. Bibcode:2010JVPal..30..163A. doi:10.1080/02724630903409220.
Bennett SC (2000). "Pterosaur flight: the role of actinofibrils in wing function". Historical Biology. 14 (4): 255–84. Bibcode:2000HBio...14..255B. doi:10.1080/10292380009380572. S2CID 85185457.
Claessens LP, O'Connor PM, Unwin DM (2009). Sereno P (ed.). "Respiratory evolution facilitated the origin of pterosaur flight and aerial gigantism". PLOS ONE. 4 (2) e4497. Bibcode:2009PLoSO...4.4497C. doi:10.1371/journal.pone.0004497. PMC 2637988. PMID 19223979.
Unwin DM, Bakhurina NN (1994). "Sordes pilosus and the nature of the pterosaur flight apparatus". Nature. 371 (6492): 62–64. Bibcode:1994Natur.371...62U. doi:10.1038/371062a0. S2CID 4314989.
Wang X, Zhou Z, Zhang F, Xu X (2002). "A nearly completely articulated rhamphorhynchoid pterosaur with exceptionally well-preserved wing membranes and "hairs" from Inner Mongolia, northeast China". Chinese Science Bulletin. 47 (3): 3. Bibcode:2002ChSBu..47..226W. doi:10.1360/02tb9054 (inactive 12 July 2025). S2CID 86641794.{{cite journal}}: CS1 maint: DOI inactive as of July 2025 (link)
Frey, E.; Tischlinger, H.; Buchy, M.-C.; Martill, D. M. (2003). "New specimens of Pterosauria (Reptilia) with soft parts with implications for pterosaurian anatomy and locomotion". Geological Society, London, Special Publications. 217 (1): 233–66. Bibcode:2003GSLSP.217..233F. doi:10.1144/GSL.SP.2003.217.01.14. S2CID 130462931.
Zhou, Chang-Fu; Schoch, Rainer R. (2011). "New material of the non-pterodactyloid pterosaur Changchengopterus pani Lü, 2009 from the Late Jurassic Tiaojishan Formation of western Liaoning". Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen. 260 (3): 265–75. Bibcode:2011NJGPA.260..265Z. doi:10.1127/0077-7749/2011/0131.
Pittman, Michael; Barlow, Luke A.; Kaye, Thomas G.; Habib, Michael B. (2021). "Pterosaurs evolved a muscular wing–body junction providing multifaceted flight performance benefits: Advanced aerodynamic smoothing, sophisticated wing root control, and wing force generation". Proceedings of the National Academy of Sciences. 118 (44) e2107631118. Bibcode:2021PNAS..11807631P. doi:10.1073/pnas.2107631118. ISSN 0027-8424. PMC 8612209. PMID 34663691. S2CID 239028043.
Unwin, David M.; Martill, David M. (December 2020). "No protofeathers on pterosaurs". Nature Ecology & Evolution. 4 (12): 1590–1591. Bibcode:2020NatEE...4.1590U. doi:10.1038/s41559-020-01308-9. PMID 32989266. S2CID 222168569.
Unwin, David M.; Bakhurina, Natasha N. (September 1994). "Sordes pilosus and the nature of the pterosaur flight apparatus". Nature. 371 (6492): 62–64. Bibcode:1994Natur.371...62U. doi:10.1038/371062a0. S2CID 4314989.
Yang, Zixiao; Jiang, Baoyu; McNamara, Maria E.; Kearns, Stuart L.; Pittman, Michael; Kaye, Thomas G.; Orr, Patrick J.; Xu, Xing; Benton, Michael J. (December 2020). "Reply to: No protofeathers on pterosaurs". Nature Ecology & Evolution. 4 (12): 1592–1593. Bibcode:2020NatEE...4.1592Y. doi:10.1038/s41559-020-01309-8. hdl:10468/11874. PMID 32989267. S2CID 222163211.
Cincotta; et al. (2022). "Pterosaur melanosomes support signalling functions for early feathers". Nature. 604 (7907): 684–688. Bibcode:2022Natur.604..684C. doi:10.1038/s41586-022-04622-3. PMC 9046085. PMID 35444275.

bbc.com (Global: 26^th place; English: 23^rd place)

Briggs, Helen (2018-12-17). "Fur flies over new pterosaur fossils". BBC News. Retrieved 2018-12-19.

biodiversitylibrary.org (Global: 168^th place; English: 136^th place)

S. Christopher Bennett (1994). "Taxonomy and systematics of the Late Cretaceous pterosaur Pteranodon (Pterosauria, Pterodactyloidea)". Occasional Papers of the Natural History Museum of the University of Kansas. 169: 1–70.
Williston, S.W. (1897). "Restoration of Ornithostoma (Pteranodon)". Kansas University Quarterly. 6: 35–51.

bris.ac.uk (Global: 5,759^th place; English: 4,110^th place)

research-information.bris.ac.uk

Yang, Zixiao; Jiang, Baoyu; McNamara, Maria E.; Kearns, Stuart L.; Pittman, Michael; Kaye, Thomas G.; Orr, Patrick J.; Xu, Xing; Benton, Michael J. (January 2019). "Pterosaur integumentary structures with complex feather-like branching". Nature Ecology & Evolution. 3 (1): 24–30. doi:10.1038/s41559-018-0728-7. hdl:1983/1f7893a1-924d-4cb3-a4bf-c4b1592356e9. PMID 30568282. S2CID 56480710.

doi.org (Global: 2^nd place; English: 2^nd place)

Witmer LM, Chatterjee S, Franzosa J, Rowe T (2003). "Neuroanatomy of flying reptiles and implications for flight, posture and behaviour" (PDF). Nature. 425 (6961): 950–53. Bibcode:2003Natur.425..950W. doi:10.1038/nature02048. PMID 14586467. S2CID 4431861.
Hone, D. W. E.; Benton, M. J. (2007). "Cope's Rule in the Pterosauria, and differing perceptions of Cope's Rule at different taxonomic levels". Journal of Evolutionary Biology. 20 (3): 1164–1170. doi:10.1111/j.1420-9101.2006.01284.x.
Benson, R. B. J.; Frigot, R. A.; Goswami, A.; Andres, B.; Bulter, R. J. (2014). "Competition and constraint drove Cope's rule in the evolution of giant flying reptiles". Nature Communications. 5 (1): 3567. doi:10.1038/ncomms4567.
Smith, RE; Chinsamy, A; Unwin, DM; Ibrahim, N; Zouhri, S; Martill, DM (2022). "Small, immature pterosaurs from the Cretaceous of Africa: implications for taphonomic bias and palaeocommunity structure in flying reptiles". Cretaceous Research. 130 105061. doi:10.1016/j.cretres.2021.105061. S2CID 239257717.
Etienne, J. L.; Smith, R. E.; Unwin, D. M.; Smyth, R. S. H.; Martill, D. M. (2024). "A 'giant' pterodactyloid pterosaur from the British Jurassic". Proceedings of the Geologists' Association. 135 (3): 335–348. Bibcode:2024PrGA..135..335E. doi:10.1016/j.pgeola.2024.05.002.
Jagielska, N.; Challands, T. J.; O'Sullivan, M.; Ross, D. A.; Fraser, N. C.; Wilkinson, M.; Brusatte, S. L. (2023). "New postcranial remains from the Lealt Shale Formation of the Isle of Skye, Scotland, showcase hidden pterosaur diversity in the Middle Jurassic". Scottish Journal of Geology. 59 (1–2): 001. Bibcode:2023ScJG...59....1J. doi:10.1144/sjg2023-001. hdl:20.500.11820/8bc004a4-ab80-4f9f-965d-f211f18e9876. S2CID 258232744.
Martin-Silverstone, Elizabeth; Witton, Mark P.; Arbour, Victoria M.; Currie, Philip J. (2016). "A small azhdarchoid pterosaur from the latest Cretaceous, the age of flying giants". Royal Society Open Science. 3 (8) 160333. Bibcode:2016RSOS....360333M. doi:10.1098/rsos.160333. PMC 5108964. PMID 27853614.
Hone, D.W.E. (2020). "A review of the taxonomy and palaeoecology of the Anurognathidae (Reptilia, Pterosauria)". Acta Geologica Sinica - English Edition. 94 (5): 1676–1692. Bibcode:2020AcGlS..94.1676H. doi:10.1111/1755-6724.14585.
Naish, D; Witton, MP; Martin-Silverstone, E (2021). "Powered flight in hatchling pterosaurs: evidence from wing form and bone strength". Scientific Reports. 11 (1) 13130. Bibcode:2021NatSR..1113130N. doi:10.1038/s41598-021-92499-z. PMC 8298463. PMID 34294737.
Witton, M.P.; Habib, M.B. (2010). "On the Size and Flight Diversity of Giant Pterosaurs, the Use of Birds as Pterosaur Analogues and Comments on Pterosaur Flightlessness". PLOS ONE. 5 (11) e13982. Bibcode:2010PLoSO...513982W. doi:10.1371/journal.pone.0013982. PMC 2981443. PMID 21085624.
Andres, B.; Langston, W. Jr. (2021). "Morphology and taxonomy of Quetzalcoatlus Lawson 1975 (Pterodactyloidea: Azhdarchoidea)". Journal of Vertebrate Paleontology. 41 (sup1): 46–202. Bibcode:2021JVPal..41S..46A. doi:10.1080/02724634.2021.1907587. ISSN 0272-4634. S2CID 245125409.
Frey E, Martill DM (1998). "Soft tissue preservation in a specimen of Pterodactylus kochi (Wagner) from the Upper Jurassic of Germany". Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen. 210 (3): 421–41. Bibcode:1998NJGPA.210..421F. doi:10.1127/njgpa/210/1998/421.
Andres, B.; Ji Q. (2008). "A new pterosaur from the Liaoning Province of China, the phylogeny of the Pterodactyloidea, and convergence in their cervical vertebrae". Palaeontology. 51 (2): 453–469. Bibcode:2008Palgy..51..453A. doi:10.1111/j.1475-4983.2008.00761.x.
Andres, B.; Clark, J. M.; Xing, X. (2010). "A new rhamphorhynchid pterosaur from the Upper Jurassic of Xinjiang, China, and the phylogenetic relationships of basal pterosaurs" (PDF). Journal of Vertebrate Paleontology. 30 (1): 163–187. Bibcode:2010JVPal..30..163A. doi:10.1080/02724630903409220.
Bennett SC (2000). "Pterosaur flight: the role of actinofibrils in wing function". Historical Biology. 14 (4): 255–84. Bibcode:2000HBio...14..255B. doi:10.1080/10292380009380572. S2CID 85185457.
Kellner, A.W.A.; Wang, X.; Tischlinger, H.; Campos, D.; Hone, D.W.E.; Meng, X. (2009). "The soft tissue of Jeholopterus (Pterosauria, Anurognathidae, Batrachognathinae) and the structure of the pterosaur wing membrane". Proceedings of the Royal Society B. 277 (1679): 321–29. doi:10.1098/rspb.2009.0846. PMC 2842671. PMID 19656798.
Claessens LP, O'Connor PM, Unwin DM (2009). Sereno P (ed.). "Respiratory evolution facilitated the origin of pterosaur flight and aerial gigantism". PLOS ONE. 4 (2) e4497. Bibcode:2009PLoSO...4.4497C. doi:10.1371/journal.pone.0004497. PMC 2637988. PMID 19223979.
Unwin DM, Bakhurina NN (1994). "Sordes pilosus and the nature of the pterosaur flight apparatus". Nature. 371 (6492): 62–64. Bibcode:1994Natur.371...62U. doi:10.1038/371062a0. S2CID 4314989.
Wang X, Zhou Z, Zhang F, Xu X (2002). "A nearly completely articulated rhamphorhynchoid pterosaur with exceptionally well-preserved wing membranes and "hairs" from Inner Mongolia, northeast China". Chinese Science Bulletin. 47 (3): 3. Bibcode:2002ChSBu..47..226W. doi:10.1360/02tb9054 (inactive 12 July 2025). S2CID 86641794.{{cite journal}}: CS1 maint: DOI inactive as of July 2025 (link)
Frey, E.; Tischlinger, H.; Buchy, M.-C.; Martill, D. M. (2003). "New specimens of Pterosauria (Reptilia) with soft parts with implications for pterosaurian anatomy and locomotion". Geological Society, London, Special Publications. 217 (1): 233–66. Bibcode:2003GSLSP.217..233F. doi:10.1144/GSL.SP.2003.217.01.14. S2CID 130462931.
Dyke GJ, Nudds RL, Rayner JM (July 2006). "Limb disparity and wing shape in pterosaurs". J. Evol. Biol. 19 (4): 1339–42. doi:10.1111/j.1420-9101.2006.01096.x. PMID 16780534. S2CID 30516133.
Wilkinson MT, Unwin DM, Ellington CP (2006). "High lift function of the pteroid bone and forewing of pterosaurs". Proceedings of the Royal Society B. 273 (1582): 119–26. doi:10.1098/rspb.2005.3278. PMC 1560000. PMID 16519243.
Bennett SC (2007). "Articulation and Function of the Pteroid Bone of Pterosaurs" (PDF). Journal of Vertebrate Paleontology. 27 (4): 881–91. doi:10.1671/0272-4634(2007)27[881:AAFOTP]2.0.CO;2. S2CID 86326537.
Zhou, Chang-Fu; Schoch, Rainer R. (2011). "New material of the non-pterodactyloid pterosaur Changchengopterus pani Lü, 2009 from the Late Jurassic Tiaojishan Formation of western Liaoning". Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen. 260 (3): 265–75. Bibcode:2011NJGPA.260..265Z. doi:10.1127/0077-7749/2011/0131.
Wang, Xiao-Lin; Kellner, Alexander W. A.; Jiang, Shun-Xing; Cheng, Xin; Meng, Xi; Rodrigues, Taissa (2010). "New long-tailed pterosaurs (Wukongopteridae) from western Liaoning, China". Anais da Academia Brasileira de Ciências. 82 (4): 1045–62. doi:10.1590/s0001-37652010000400024. PMID 21152776.
Wilkinson M.T.; Unwin D.M.; Ellington C.P. (2006). "High lift function of the pteroid bone and forewing of pterosaurs". Proceedings of the Royal Society B. 273 (1582): 119–26. doi:10.1098/rspb.2005.3278. PMC 1560000. PMID 16519243.
Pittman, Michael; Barlow, Luke A.; Kaye, Thomas G.; Habib, Michael B. (2021). "Pterosaurs evolved a muscular wing–body junction providing multifaceted flight performance benefits: Advanced aerodynamic smoothing, sophisticated wing root control, and wing force generation". Proceedings of the National Academy of Sciences. 118 (44) e2107631118. Bibcode:2021PNAS..11807631P. doi:10.1073/pnas.2107631118. ISSN 0027-8424. PMC 8612209. PMID 34663691. S2CID 239028043.
Yang, Zixiao; Jiang, Baoyu; McNamara, Maria E.; Kearns, Stuart L.; Pittman, Michael; Kaye, Thomas G.; Orr, Patrick J.; Xu, Xing; Benton, Michael J. (January 2019). "Pterosaur integumentary structures with complex feather-like branching". Nature Ecology & Evolution. 3 (1): 24–30. doi:10.1038/s41559-018-0728-7. hdl:1983/1f7893a1-924d-4cb3-a4bf-c4b1592356e9. PMID 30568282. S2CID 56480710.
Unwin, David M.; Martill, David M. (December 2020). "No protofeathers on pterosaurs". Nature Ecology & Evolution. 4 (12): 1590–1591. Bibcode:2020NatEE...4.1590U. doi:10.1038/s41559-020-01308-9. PMID 32989266. S2CID 222168569.
Kellner; et al. (2009). "The Soft Tissue of Jeholopterus (Pterosauria, Anurognathidae, Batrachognathinae) and the Structure of the Pterosaur Wing Membrane". Proceedings of the Royal Society B: Biological Sciences. 277 (1679): 321–29. doi:10.1098/rspb.2009.0846. PMC 2842671. PMID 19656798.
Unwin, David M.; Bakhurina, Natasha N. (September 1994). "Sordes pilosus and the nature of the pterosaur flight apparatus". Nature. 371 (6492): 62–64. Bibcode:1994Natur.371...62U. doi:10.1038/371062a0. S2CID 4314989.
Yang, Zixiao; Jiang, Baoyu; McNamara, Maria E.; Kearns, Stuart L.; Pittman, Michael; Kaye, Thomas G.; Orr, Patrick J.; Xu, Xing; Benton, Michael J. (December 2020). "Reply to: No protofeathers on pterosaurs". Nature Ecology & Evolution. 4 (12): 1592–1593. Bibcode:2020NatEE...4.1592Y. doi:10.1038/s41559-020-01309-8. hdl:10468/11874. PMID 32989267. S2CID 222163211.
D'Alba, Liliana (January 2019). "Pterosaur plumage". Nature Ecology & Evolution. 3 (1): 12–13. doi:10.1038/s41559-018-0767-0. PMID 30568284. S2CID 56480834.
Cincotta; et al. (2022). "Pterosaur melanosomes support signalling functions for early feathers". Nature. 604 (7907): 684–688. Bibcode:2022Natur.604..684C. doi:10.1038/s41586-022-04622-3. PMC 9046085. PMID 35444275.

hdl.handle.net (Global: 66^th place; English: 34^th place)

Jagielska, N.; Challands, T. J.; O'Sullivan, M.; Ross, D. A.; Fraser, N. C.; Wilkinson, M.; Brusatte, S. L. (2023). "New postcranial remains from the Lealt Shale Formation of the Isle of Skye, Scotland, showcase hidden pterosaur diversity in the Middle Jurassic". Scottish Journal of Geology. 59 (1–2): 001. Bibcode:2023ScJG...59....1J. doi:10.1144/sjg2023-001. hdl:20.500.11820/8bc004a4-ab80-4f9f-965d-f211f18e9876. S2CID 258232744.
Yang, Zixiao; Jiang, Baoyu; McNamara, Maria E.; Kearns, Stuart L.; Pittman, Michael; Kaye, Thomas G.; Orr, Patrick J.; Xu, Xing; Benton, Michael J. (January 2019). "Pterosaur integumentary structures with complex feather-like branching". Nature Ecology & Evolution. 3 (1): 24–30. doi:10.1038/s41559-018-0728-7. hdl:1983/1f7893a1-924d-4cb3-a4bf-c4b1592356e9. PMID 30568282. S2CID 56480710.
Yang, Zixiao; Jiang, Baoyu; McNamara, Maria E.; Kearns, Stuart L.; Pittman, Michael; Kaye, Thomas G.; Orr, Patrick J.; Xu, Xing; Benton, Michael J. (December 2020). "Reply to: No protofeathers on pterosaurs". Nature Ecology & Evolution. 4 (12): 1592–1593. Bibcode:2020NatEE...4.1592Y. doi:10.1038/s41559-020-01309-8. hdl:10468/11874. PMID 32989267. S2CID 222163211.

nature.com (Global: 207^th place; English: 347^th place)

Benson, R. B. J.; Frigot, R. A.; Goswami, A.; Andres, B.; Bulter, R. J. (2014). "Competition and constraint drove Cope's rule in the evolution of giant flying reptiles". Nature Communications. 5 (1): 3567. doi:10.1038/ncomms4567.

nih.gov (Global: 5^th place; English: 5^th place)

pubmed.ncbi.nlm.nih.gov

Witmer LM, Chatterjee S, Franzosa J, Rowe T (2003). "Neuroanatomy of flying reptiles and implications for flight, posture and behaviour" (PDF). Nature. 425 (6961): 950–53. Bibcode:2003Natur.425..950W. doi:10.1038/nature02048. PMID 14586467. S2CID 4431861.
Martin-Silverstone, Elizabeth; Witton, Mark P.; Arbour, Victoria M.; Currie, Philip J. (2016). "A small azhdarchoid pterosaur from the latest Cretaceous, the age of flying giants". Royal Society Open Science. 3 (8) 160333. Bibcode:2016RSOS....360333M. doi:10.1098/rsos.160333. PMC 5108964. PMID 27853614.
Naish, D; Witton, MP; Martin-Silverstone, E (2021). "Powered flight in hatchling pterosaurs: evidence from wing form and bone strength". Scientific Reports. 11 (1) 13130. Bibcode:2021NatSR..1113130N. doi:10.1038/s41598-021-92499-z. PMC 8298463. PMID 34294737.
Witton, M.P.; Habib, M.B. (2010). "On the Size and Flight Diversity of Giant Pterosaurs, the Use of Birds as Pterosaur Analogues and Comments on Pterosaur Flightlessness". PLOS ONE. 5 (11) e13982. Bibcode:2010PLoSO...513982W. doi:10.1371/journal.pone.0013982. PMC 2981443. PMID 21085624.
Kellner, A.W.A.; Wang, X.; Tischlinger, H.; Campos, D.; Hone, D.W.E.; Meng, X. (2009). "The soft tissue of Jeholopterus (Pterosauria, Anurognathidae, Batrachognathinae) and the structure of the pterosaur wing membrane". Proceedings of the Royal Society B. 277 (1679): 321–29. doi:10.1098/rspb.2009.0846. PMC 2842671. PMID 19656798.
Claessens LP, O'Connor PM, Unwin DM (2009). Sereno P (ed.). "Respiratory evolution facilitated the origin of pterosaur flight and aerial gigantism". PLOS ONE. 4 (2) e4497. Bibcode:2009PLoSO...4.4497C. doi:10.1371/journal.pone.0004497. PMC 2637988. PMID 19223979.
Dyke GJ, Nudds RL, Rayner JM (July 2006). "Limb disparity and wing shape in pterosaurs". J. Evol. Biol. 19 (4): 1339–42. doi:10.1111/j.1420-9101.2006.01096.x. PMID 16780534. S2CID 30516133.
Wilkinson MT, Unwin DM, Ellington CP (2006). "High lift function of the pteroid bone and forewing of pterosaurs". Proceedings of the Royal Society B. 273 (1582): 119–26. doi:10.1098/rspb.2005.3278. PMC 1560000. PMID 16519243.
Wang, Xiao-Lin; Kellner, Alexander W. A.; Jiang, Shun-Xing; Cheng, Xin; Meng, Xi; Rodrigues, Taissa (2010). "New long-tailed pterosaurs (Wukongopteridae) from western Liaoning, China". Anais da Academia Brasileira de Ciências. 82 (4): 1045–62. doi:10.1590/s0001-37652010000400024. PMID 21152776.
Wilkinson M.T.; Unwin D.M.; Ellington C.P. (2006). "High lift function of the pteroid bone and forewing of pterosaurs". Proceedings of the Royal Society B. 273 (1582): 119–26. doi:10.1098/rspb.2005.3278. PMC 1560000. PMID 16519243.
Pittman, Michael; Barlow, Luke A.; Kaye, Thomas G.; Habib, Michael B. (2021). "Pterosaurs evolved a muscular wing–body junction providing multifaceted flight performance benefits: Advanced aerodynamic smoothing, sophisticated wing root control, and wing force generation". Proceedings of the National Academy of Sciences. 118 (44) e2107631118. Bibcode:2021PNAS..11807631P. doi:10.1073/pnas.2107631118. ISSN 0027-8424. PMC 8612209. PMID 34663691. S2CID 239028043.
Yang, Zixiao; Jiang, Baoyu; McNamara, Maria E.; Kearns, Stuart L.; Pittman, Michael; Kaye, Thomas G.; Orr, Patrick J.; Xu, Xing; Benton, Michael J. (January 2019). "Pterosaur integumentary structures with complex feather-like branching". Nature Ecology & Evolution. 3 (1): 24–30. doi:10.1038/s41559-018-0728-7. hdl:1983/1f7893a1-924d-4cb3-a4bf-c4b1592356e9. PMID 30568282. S2CID 56480710.
Unwin, David M.; Martill, David M. (December 2020). "No protofeathers on pterosaurs". Nature Ecology & Evolution. 4 (12): 1590–1591. Bibcode:2020NatEE...4.1590U. doi:10.1038/s41559-020-01308-9. PMID 32989266. S2CID 222168569.
Kellner; et al. (2009). "The Soft Tissue of Jeholopterus (Pterosauria, Anurognathidae, Batrachognathinae) and the Structure of the Pterosaur Wing Membrane". Proceedings of the Royal Society B: Biological Sciences. 277 (1679): 321–29. doi:10.1098/rspb.2009.0846. PMC 2842671. PMID 19656798.
Yang, Zixiao; Jiang, Baoyu; McNamara, Maria E.; Kearns, Stuart L.; Pittman, Michael; Kaye, Thomas G.; Orr, Patrick J.; Xu, Xing; Benton, Michael J. (December 2020). "Reply to: No protofeathers on pterosaurs". Nature Ecology & Evolution. 4 (12): 1592–1593. Bibcode:2020NatEE...4.1592Y. doi:10.1038/s41559-020-01309-8. hdl:10468/11874. PMID 32989267. S2CID 222163211.
D'Alba, Liliana (January 2019). "Pterosaur plumage". Nature Ecology & Evolution. 3 (1): 12–13. doi:10.1038/s41559-018-0767-0. PMID 30568284. S2CID 56480834.
Cincotta; et al. (2022). "Pterosaur melanosomes support signalling functions for early feathers". Nature. 604 (7907): 684–688. Bibcode:2022Natur.604..684C. doi:10.1038/s41586-022-04622-3. PMC 9046085. PMID 35444275.

ncbi.nlm.nih.gov

Martin-Silverstone, Elizabeth; Witton, Mark P.; Arbour, Victoria M.; Currie, Philip J. (2016). "A small azhdarchoid pterosaur from the latest Cretaceous, the age of flying giants". Royal Society Open Science. 3 (8) 160333. Bibcode:2016RSOS....360333M. doi:10.1098/rsos.160333. PMC 5108964. PMID 27853614.
Naish, D; Witton, MP; Martin-Silverstone, E (2021). "Powered flight in hatchling pterosaurs: evidence from wing form and bone strength". Scientific Reports. 11 (1) 13130. Bibcode:2021NatSR..1113130N. doi:10.1038/s41598-021-92499-z. PMC 8298463. PMID 34294737.
Witton, M.P.; Habib, M.B. (2010). "On the Size and Flight Diversity of Giant Pterosaurs, the Use of Birds as Pterosaur Analogues and Comments on Pterosaur Flightlessness". PLOS ONE. 5 (11) e13982. Bibcode:2010PLoSO...513982W. doi:10.1371/journal.pone.0013982. PMC 2981443. PMID 21085624.
Kellner, A.W.A.; Wang, X.; Tischlinger, H.; Campos, D.; Hone, D.W.E.; Meng, X. (2009). "The soft tissue of Jeholopterus (Pterosauria, Anurognathidae, Batrachognathinae) and the structure of the pterosaur wing membrane". Proceedings of the Royal Society B. 277 (1679): 321–29. doi:10.1098/rspb.2009.0846. PMC 2842671. PMID 19656798.
Claessens LP, O'Connor PM, Unwin DM (2009). Sereno P (ed.). "Respiratory evolution facilitated the origin of pterosaur flight and aerial gigantism". PLOS ONE. 4 (2) e4497. Bibcode:2009PLoSO...4.4497C. doi:10.1371/journal.pone.0004497. PMC 2637988. PMID 19223979.
Wilkinson MT, Unwin DM, Ellington CP (2006). "High lift function of the pteroid bone and forewing of pterosaurs". Proceedings of the Royal Society B. 273 (1582): 119–26. doi:10.1098/rspb.2005.3278. PMC 1560000. PMID 16519243.
Wilkinson M.T.; Unwin D.M.; Ellington C.P. (2006). "High lift function of the pteroid bone and forewing of pterosaurs". Proceedings of the Royal Society B. 273 (1582): 119–26. doi:10.1098/rspb.2005.3278. PMC 1560000. PMID 16519243.
Pittman, Michael; Barlow, Luke A.; Kaye, Thomas G.; Habib, Michael B. (2021). "Pterosaurs evolved a muscular wing–body junction providing multifaceted flight performance benefits: Advanced aerodynamic smoothing, sophisticated wing root control, and wing force generation". Proceedings of the National Academy of Sciences. 118 (44) e2107631118. Bibcode:2021PNAS..11807631P. doi:10.1073/pnas.2107631118. ISSN 0027-8424. PMC 8612209. PMID 34663691. S2CID 239028043.
Kellner; et al. (2009). "The Soft Tissue of Jeholopterus (Pterosauria, Anurognathidae, Batrachognathinae) and the Structure of the Pterosaur Wing Membrane". Proceedings of the Royal Society B: Biological Sciences. 277 (1679): 321–29. doi:10.1098/rspb.2009.0846. PMC 2842671. PMID 19656798.
Cincotta; et al. (2022). "Pterosaur melanosomes support signalling functions for early feathers". Nature. 604 (7907): 684–688. Bibcode:2022Natur.604..684C. doi:10.1038/s41586-022-04622-3. PMC 9046085. PMID 35444275.

oup.com (Global: 292^nd place; English: 233^rd place)

academic.oup.com

Hone, D. W. E.; Benton, M. J. (2007). "Cope's Rule in the Pterosauria, and differing perceptions of Cope's Rule at different taxonomic levels". Journal of Evolutionary Biology. 20 (3): 1164–1170. doi:10.1111/j.1420-9101.2006.01284.x.

rero.ch (Global: 3,023^rd place; English: 2,092^nd place)

doc.rero.ch

Witmer LM, Chatterjee S, Franzosa J, Rowe T (2003). "Neuroanatomy of flying reptiles and implications for flight, posture and behaviour" (PDF). Nature. 425 (6961): 950–53. Bibcode:2003Natur.425..950W. doi:10.1038/nature02048. PMID 14586467. S2CID 4431861.
Andres, B.; Clark, J. M.; Xing, X. (2010). "A new rhamphorhynchid pterosaur from the Upper Jurassic of Xinjiang, China, and the phylogenetic relationships of basal pterosaurs" (PDF). Journal of Vertebrate Paleontology. 30 (1): 163–187. Bibcode:2010JVPal..30..163A. doi:10.1080/02724630903409220.
Bennett SC (2007). "Articulation and Function of the Pteroid Bone of Pterosaurs" (PDF). Journal of Vertebrate Paleontology. 27 (4): 881–91. doi:10.1671/0272-4634(2007)27[881:AAFOTP]2.0.CO;2. S2CID 86326537.

researchgate.net (Global: 96^th place; English: 89^th place)

Andres, B.; Ji Q. (2008). "A new pterosaur from the Liaoning Province of China, the phylogeny of the Pterodactyloidea, and convergence in their cervical vertebrae". Palaeontology. 51 (2): 453–469. Bibcode:2008Palgy..51..453A. doi:10.1111/j.1475-4983.2008.00761.x.

sciencedirect.com (Global: 137^th place; English: 146^th place)

Smith, RE; Chinsamy, A; Unwin, DM; Ibrahim, N; Zouhri, S; Martill, DM (2022). "Small, immature pterosaurs from the Cretaceous of Africa: implications for taphonomic bias and palaeocommunity structure in flying reptiles". Cretaceous Research. 130 105061. doi:10.1016/j.cretres.2021.105061. S2CID 239257717.

semanticscholar.org (Global: 15^th place; English: 8^th place)

api.semanticscholar.org

Witmer LM, Chatterjee S, Franzosa J, Rowe T (2003). "Neuroanatomy of flying reptiles and implications for flight, posture and behaviour" (PDF). Nature. 425 (6961): 950–53. Bibcode:2003Natur.425..950W. doi:10.1038/nature02048. PMID 14586467. S2CID 4431861.
Smith, RE; Chinsamy, A; Unwin, DM; Ibrahim, N; Zouhri, S; Martill, DM (2022). "Small, immature pterosaurs from the Cretaceous of Africa: implications for taphonomic bias and palaeocommunity structure in flying reptiles". Cretaceous Research. 130 105061. doi:10.1016/j.cretres.2021.105061. S2CID 239257717.
Jagielska, N.; Challands, T. J.; O'Sullivan, M.; Ross, D. A.; Fraser, N. C.; Wilkinson, M.; Brusatte, S. L. (2023). "New postcranial remains from the Lealt Shale Formation of the Isle of Skye, Scotland, showcase hidden pterosaur diversity in the Middle Jurassic". Scottish Journal of Geology. 59 (1–2): 001. Bibcode:2023ScJG...59....1J. doi:10.1144/sjg2023-001. hdl:20.500.11820/8bc004a4-ab80-4f9f-965d-f211f18e9876. S2CID 258232744.
Andres, B.; Langston, W. Jr. (2021). "Morphology and taxonomy of Quetzalcoatlus Lawson 1975 (Pterodactyloidea: Azhdarchoidea)". Journal of Vertebrate Paleontology. 41 (sup1): 46–202. Bibcode:2021JVPal..41S..46A. doi:10.1080/02724634.2021.1907587. ISSN 0272-4634. S2CID 245125409.
Bennett SC (2000). "Pterosaur flight: the role of actinofibrils in wing function". Historical Biology. 14 (4): 255–84. Bibcode:2000HBio...14..255B. doi:10.1080/10292380009380572. S2CID 85185457.
Unwin DM, Bakhurina NN (1994). "Sordes pilosus and the nature of the pterosaur flight apparatus". Nature. 371 (6492): 62–64. Bibcode:1994Natur.371...62U. doi:10.1038/371062a0. S2CID 4314989.
Wang X, Zhou Z, Zhang F, Xu X (2002). "A nearly completely articulated rhamphorhynchoid pterosaur with exceptionally well-preserved wing membranes and "hairs" from Inner Mongolia, northeast China". Chinese Science Bulletin. 47 (3): 3. Bibcode:2002ChSBu..47..226W. doi:10.1360/02tb9054 (inactive 12 July 2025). S2CID 86641794.{{cite journal}}: CS1 maint: DOI inactive as of July 2025 (link)
Frey, E.; Tischlinger, H.; Buchy, M.-C.; Martill, D. M. (2003). "New specimens of Pterosauria (Reptilia) with soft parts with implications for pterosaurian anatomy and locomotion". Geological Society, London, Special Publications. 217 (1): 233–66. Bibcode:2003GSLSP.217..233F. doi:10.1144/GSL.SP.2003.217.01.14. S2CID 130462931.
Dyke GJ, Nudds RL, Rayner JM (July 2006). "Limb disparity and wing shape in pterosaurs". J. Evol. Biol. 19 (4): 1339–42. doi:10.1111/j.1420-9101.2006.01096.x. PMID 16780534. S2CID 30516133.
Bennett SC (2007). "Articulation and Function of the Pteroid Bone of Pterosaurs" (PDF). Journal of Vertebrate Paleontology. 27 (4): 881–91. doi:10.1671/0272-4634(2007)27[881:AAFOTP]2.0.CO;2. S2CID 86326537.
Pittman, Michael; Barlow, Luke A.; Kaye, Thomas G.; Habib, Michael B. (2021). "Pterosaurs evolved a muscular wing–body junction providing multifaceted flight performance benefits: Advanced aerodynamic smoothing, sophisticated wing root control, and wing force generation". Proceedings of the National Academy of Sciences. 118 (44) e2107631118. Bibcode:2021PNAS..11807631P. doi:10.1073/pnas.2107631118. ISSN 0027-8424. PMC 8612209. PMID 34663691. S2CID 239028043.
Yang, Zixiao; Jiang, Baoyu; McNamara, Maria E.; Kearns, Stuart L.; Pittman, Michael; Kaye, Thomas G.; Orr, Patrick J.; Xu, Xing; Benton, Michael J. (January 2019). "Pterosaur integumentary structures with complex feather-like branching". Nature Ecology & Evolution. 3 (1): 24–30. doi:10.1038/s41559-018-0728-7. hdl:1983/1f7893a1-924d-4cb3-a4bf-c4b1592356e9. PMID 30568282. S2CID 56480710.
Unwin, David M.; Martill, David M. (December 2020). "No protofeathers on pterosaurs". Nature Ecology & Evolution. 4 (12): 1590–1591. Bibcode:2020NatEE...4.1590U. doi:10.1038/s41559-020-01308-9. PMID 32989266. S2CID 222168569.
Unwin, David M.; Bakhurina, Natasha N. (September 1994). "Sordes pilosus and the nature of the pterosaur flight apparatus". Nature. 371 (6492): 62–64. Bibcode:1994Natur.371...62U. doi:10.1038/371062a0. S2CID 4314989.
Yang, Zixiao; Jiang, Baoyu; McNamara, Maria E.; Kearns, Stuart L.; Pittman, Michael; Kaye, Thomas G.; Orr, Patrick J.; Xu, Xing; Benton, Michael J. (December 2020). "Reply to: No protofeathers on pterosaurs". Nature Ecology & Evolution. 4 (12): 1592–1593. Bibcode:2020NatEE...4.1592Y. doi:10.1038/s41559-020-01309-8. hdl:10468/11874. PMID 32989267. S2CID 222163211.
D'Alba, Liliana (January 2019). "Pterosaur plumage". Nature Ecology & Evolution. 3 (1): 12–13. doi:10.1038/s41559-018-0767-0. PMID 30568284. S2CID 56480834.

worldcat.org (Global: 4^th place; English: 4^th place)

search.worldcat.org

Andres, B.; Langston, W. Jr. (2021). "Morphology and taxonomy of Quetzalcoatlus Lawson 1975 (Pterodactyloidea: Azhdarchoidea)". Journal of Vertebrate Paleontology. 41 (sup1): 46–202. Bibcode:2021JVPal..41S..46A. doi:10.1080/02724634.2021.1907587. ISSN 0272-4634. S2CID 245125409.
Pittman, Michael; Barlow, Luke A.; Kaye, Thomas G.; Habib, Michael B. (2021). "Pterosaurs evolved a muscular wing–body junction providing multifaceted flight performance benefits: Advanced aerodynamic smoothing, sophisticated wing root control, and wing force generation". Proceedings of the National Academy of Sciences. 118 (44) e2107631118. Bibcode:2021PNAS..11807631P. doi:10.1073/pnas.2107631118. ISSN 0027-8424. PMC 8612209. PMID 34663691. S2CID 239028043.

Anatomy of pterosaurs (English Wikipedia)

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