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Burchill, A. T.; Moreau, C. S. (5 February 2016). "Colony size evolution in ants: macroevolutionary trends". Insectes Sociaux. 63 (2): 291–298. doi:10.1007/s00040-016-0465-3. S2CID4817407.
Peeters, Christian (1 October 1991). "The occurrence of sexual reproduction among ant workers". Biological Journal of the Linnean Society. 44 (2): 141–152. doi:10.1111/j.1095-8312.1991.tb00612.x. ISSN0024-4066.