Krell, Frank-Thorsten; Krell-Westerwalbesloh, Sylvia (2024). "One elephant may sustain 2 million dung beetles in East African savannas on any given day". The Science of Nature. 111 (1). doi:10.1007/s00114-024-01894-9. ISSN0028-1042.
Skelley, P. E. Aphodiinae.Archived 2018-02-14 at the Wayback Machine Generic Guide to New World Scarab Beetles. University of Nebraska State Museum. 2008 Version.
web.archive.org
Skelley, P. E. Aphodiinae.Archived 2018-02-14 at the Wayback Machine Generic Guide to New World Scarab Beetles. University of Nebraska State Museum. 2008 Version.
worldcat.org
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Krell, Frank-Thorsten; Krell-Westerwalbesloh, Sylvia (2024). "One elephant may sustain 2 million dung beetles in East African savannas on any given day". The Science of Nature. 111 (1). doi:10.1007/s00114-024-01894-9. ISSN0028-1042.