G. David Johnson (1983). "Niphon spinosus: A Primitive Epinepheline Serranid, with Comments on the Monophyly and Intrarelationships of the Serranidae". Copeia. 777 (3): 777–787. doi:10.2307/1444346. JSTOR1444346.
G. David Johnson (1983). "Niphon spinosus: A Primitive Epinepheline Serranid, with Comments on the Monophyly and Intrarelationships of the Serranidae". Copeia. 777 (3): 777–787. doi:10.2307/1444346. JSTOR1444346.