Laura Lowe Forrest, Mark Hughes & Peter M. Hollingsworth (2005). "A phylogeny of Begonia using nuclear ribosomal sequence data and morphological characters". Systematic Botany. 30 (3): 671–682. doi:10.1600/0363644054782297. S2CID85909059.
Laura Lowe Forrest, Mark Hughes & Peter M. Hollingsworth (2005). "A phylogeny of Begonia using nuclear ribosomal sequence data and morphological characters". Systematic Botany. 30 (3): 671–682. doi:10.1600/0363644054782297. S2CID85909059.