Dubost, G. (1984), "Comparison of the diets of frugivorous forest mammals of Gabon", Journal of Mammalogy, 65 (2), American Society of Mammalogists: 298–316, doi:10.2307/1381169, JSTOR1381169
Beentje, Henk; Smith, Sara (2001), "Plant Systematics and Phytogeography for the Understanding of African Biodiversity", Systematics and Geography of Plants, 71 (2), National Botanic Garden of Belgium: 284–286, doi:10.2307/3668673, JSTOR3668673.
Dubost, G. (1984), "Comparison of the diets of frugivorous forest mammals of Gabon", Journal of Mammalogy, 65 (2), American Society of Mammalogists: 298–316, doi:10.2307/1381169, JSTOR1381169
Beentje, Henk; Smith, Sara (2001), "Plant Systematics and Phytogeography for the Understanding of African Biodiversity", Systematics and Geography of Plants, 71 (2), National Botanic Garden of Belgium: 284–286, doi:10.2307/3668673, JSTOR3668673.