Taberlet, P.; Bouvet, J. (1994-03-22). "Mitochondrial DNA Polymorphism, Phylogeography, and Conservation Genetics of the Brown Bear Ursus arctos in Europe". Proceedings of the Royal Society of London. Series B: Biological Sciences. 255 (1344): 195–200. doi:10.1098/rspb.1994.0028. ISSN0962-8452. PMID8022838.
Linnell, John D. C.; Steuer, Daniel; Odden, John; Kaczensky, Petra; Swenson, Jon E. (2002). European Brown Bear Compendium(PDF). Wildlife Conservation Issues–Technical Series (Report).
Taberlet, P.; Bouvet, J. (1994-03-22). "Mitochondrial DNA Polymorphism, Phylogeography, and Conservation Genetics of the Brown Bear Ursus arctos in Europe". Proceedings of the Royal Society of London. Series B: Biological Sciences. 255 (1344): 195–200. doi:10.1098/rspb.1994.0028. ISSN0962-8452. PMID8022838.
Taberlet, P.; Bouvet, J. (1994-03-22). "Mitochondrial DNA Polymorphism, Phylogeography, and Conservation Genetics of the Brown Bear Ursus arctos in Europe". Proceedings of the Royal Society of London. Series B: Biological Sciences. 255 (1344): 195–200. doi:10.1098/rspb.1994.0028. ISSN0962-8452. PMID8022838.