Kurochkin, E. N.; Saveliev, S. V.; Postnov, A. A.; Pervushov, E. M.; Popov, E. V. (2006). "On the brain of a primitive bird from the Upper Cretaceous of European Russia". Paleontological Journal. 40 (6): 655–667. doi:10.1134/s0031030106060086.
Walsh, S. A.; Milne, A. C. (2011). "Evolution of the avian brain and senses". In Dyke, G.; Kaiser, G. (eds.). Living Dinosaurs: The Evolutionary History of Modern Birds. Chichester: John Wiley & Sons. pp. 282–305. doi:10.1002/9781119990475.ch11. ISBN9780470656662.