Campbell, Jonathan A. & Savage, Jay M. (2000). "Taxonomic reconsideration of Middle American frogs of the Eleutherodactylus rugulosus group (Anura: Leptodactylidae): a reconnaissance of subtle nuances among frogs". Herpetological Monographs. 14: 186–292. doi:10.2307/1467048. JSTOR1467048.
Campbell, Jonathan A. & Savage, Jay M. (2000). "Taxonomic reconsideration of Middle American frogs of the Eleutherodactylus rugulosus group (Anura: Leptodactylidae): a reconnaissance of subtle nuances among frogs". Herpetological Monographs. 14: 186–292. doi:10.2307/1467048. JSTOR1467048.