Brown RM, Su YC, Barger B, Siler CD, Sanguila MB, Diesmos AC, Blackburn DC (January 2016). "Phylogeny of the island archipelago frog genus Sanguirana: Another endemic Philippine radiation that diversified 'Out-of-Palawan'". Molecular Phylogenetics and Evolution. 94 (Pt B): 531–536. doi:10.1016/j.ympev.2015.10.010. PMID26477738.
Brown RM, Su YC, Barger B, Siler CD, Sanguila MB, Diesmos AC, Blackburn DC (January 2016). "Phylogeny of the island archipelago frog genus Sanguirana: Another endemic Philippine radiation that diversified 'Out-of-Palawan'". Molecular Phylogenetics and Evolution. 94 (Pt B): 531–536. doi:10.1016/j.ympev.2015.10.010. PMID26477738.