D.M.S. Watson, "Adaptation," Nature, Vol. 124, 10 August 1929, 233. The article also appears in the Report of the Ninety-Seventh Meeting British Association for the Advancement of Science (Office of the British Association: London, 1929), 88-99 and can be accessed at https://archive.org/details/reportofbritisha30adva . The quote discussed here is on page 95.