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Matsui, M.; Hamidy, A.; Murphy, R. W.; Khonsue, W.; Yambun, P.; Shimada, T.; Ahmad, N.; Belabut, D. M.; Jiang, J. P. (2010). "Phylogenetic relationships of megophryid frogs of the genus Leptobrachium (Amphibia, Anura) as revealed by mtDNA gene sequences". Molecular Phylogenetics and Evolution. 56 (1): 259–272. doi:10.1016/j.ympev.2010.03.014. PMID20302957.
Matsui, M.; Hamidy, A.; Murphy, R. W.; Khonsue, W.; Yambun, P.; Shimada, T.; Ahmad, N.; Belabut, D. M.; Jiang, J. P. (2010). "Phylogenetic relationships of megophryid frogs of the genus Leptobrachium (Amphibia, Anura) as revealed by mtDNA gene sequences". Molecular Phylogenetics and Evolution. 56 (1): 259–272. doi:10.1016/j.ympev.2010.03.014. PMID20302957.