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Ron, Santiago R.; Santos, Juan C. & Cannatella, David C. (2006). "Phylogeny of the túngara frog genus Engystomops (=Physalaemus pustulosus species group; Anura: Leptodactylidae)". Molecular Phylogenetics and Evolution. 39 (2): 392–403. doi:10.1016/j.ympev.2005.11.022. PMID16446105.
semanticscholar.org
api.semanticscholar.org
Narváez, Andrea E. & Ron, Santiago R. (2013). "Feeding habits of Engystomops pustulatus (Anura: Leptodactylidae) in Western Ecuador". South American Journal of Herpetology. 8 (3): 161–167. doi:10.2994/sajh-d-13-00012.1. S2CID86350285.