Selden, Paul A.; Shcherbakov, Dmitry E.; Dunlop, Jason A. & Eskov, Kirill Yu. (2013), "Arachnids from the Carboniferous of Russia and Ukraine, and the Permian of Kazakhstan", Paläontologische Zeitschrift, 88 (3): 297–307, doi:10.1007/s12542-013-0198-9, S2CID55260690
Selden, Paul A.; Shcherbakov, Dmitry E.; Dunlop, Jason A. & Eskov, Kirill Yu. (2013), "Arachnids from the Carboniferous of Russia and Ukraine, and the Permian of Kazakhstan", Paläontologische Zeitschrift, 88 (3): 297–307, doi:10.1007/s12542-013-0198-9, S2CID55260690