References analysis of the Wikipedia article "Equine coat color genetics" in the English version

doi.org

Pruvost, Melanie; Bellone, Rebecca; Benecke, Norbert; Sandoval-Castellanos, Edson; Cieslak, Michael; Kuznetsova, Tatyana; Morales-Muñiz, Arturo; O'Connor, Terry; Reissmann, Monika; Hofreiter, Michael; Ludwig, Arne (15 November 2011). "Genotypes of predomestic horses match phenotypes painted in Paleolithic works of cave art". Proceedings of the National Academy of Sciences. 108 (46): 18626–18630. Bibcode:2011PNAS..10818626P. doi:10.1073/pnas.1108982108. PMC 3219153. PMID 22065780.

Sponenberg, D. Phillip; Bellone, Rebecca (2017). "Introduction". Equine Color Genetics (4th ed.). John Wiley & Sons. pp. 1–21, 26. doi:10.1002/9781119130628.ch1. ISBN 978-1-119-13062-8.

Imsland, Freyja; McGowan, Kelly; Rubin, Carl-Johan; Henegar, Corneliu; Sundström, Elisabeth; Berglund, Jonas; Schwochow, Doreen; Gustafson, Ulla; Imsland, Páll; Lindblad-Toh, Kerstin; Lindgren, Gabriella; Mikko, Sofia; Millon, Lee; Wade, Claire; Schubert, Mikkel; Orlando, Ludovic; Penedo, Maria Cecilia T; Barsh, Gregory S; Andersson, Leif (February 2016). "Regulatory mutations in TBX3 disrupt asymmetric hair pigmentation that underlies Dun camouflage color in horses". Nature Genetics. 48 (2): 152–158. doi:10.1038/ng.3475. PMC 4731265. PMID 26691985.

"A horse of a different color: Genetics of camouflage and the dun pattern". ScienceDaily (Press release). December 21, 2015.
Ludwig, Arne; Pruvost, Melanie; Reissmann, Monika; Benecke, Norbert; Brockmann, Gudrun A.; Castaños, Pedro; Cieslak, Michael; Lippold, Sebastian; Llorente, Laura; Malaspinas, Anna-Sapfo; Stalkin, Montgomery; Hofreiter, Michael (2009). "Coat Color Variation at the Beginning of Horse Domestication". Science. 324 (5926): 485. Bibcode:2009Sci...324..485L. doi:10.1126/science.1172750. PMC 5102060. PMID 19390039.
Wutke, Saskia; Benecke, Norbert; Sandoval-Castellanos, Edson; Döhle, Hans-Jürgen; Friederich, Susanne; Gonzalez, Javier; Hallsson, Jón Hallsteinn; Hofreiter, Michael; Lõugas, Lembi; Magnell, Ola; Morales-Muniz, Arturo; Orlando, Ludovic; Pálsdóttir, Albína Hulda; Reissmann, Monika; Ruttkay, Matej; Trinks, Alexandra; Ludwig, Arne (23 December 2016). "Spotted phenotypes in horses lost attractiveness in the Middle Ages". Scientific Reports. 6 (1): 38548. Bibcode:2016NatSR...638548W. doi:10.1038/srep38548. PMC 5141471. PMID 27924839.
Rosengren Pielberg, Gerli; Golovko, Anna; Sundström, Elisabeth; Curik, Ino; Lennartsson, Johan; Seltenhammer, Monika H; Druml, Thomas; Binns, Matthew; Fitzsimmons, Carolyn; Lindgren, Gabriella; Sandberg, Kaj; Baumung, Roswitha; Vetterlein, Monika; Strömberg, Sara; Grabherr, Manfred; Wade, Claire; Lindblad-Toh, Kerstin; Pontén, Fredrik; Heldin, Carl-Henrik; Sölkner, Johann; Andersson, Leif (August 2008). "A cis-acting regulatory mutation causes premature hair graying and susceptibility to melanoma in the horse". Nature Genetics. 40 (8): 1004–1009. doi:10.1038/ng.185. PMID 18641652. S2CID 6666394.
- "Genetics Of White Horses Unraveled: One Mutation Makes Ordinary Horses Turn Grey, Then White, Very Young". ScienceDaily (Press release). July 23, 2008.
- Micillo, Raffaella; Panzella, Lucia; Koike, Kenzo; Monfrecola, Giuseppe; Napolitano, Alessandra; d'Ischia, Marco (17 May 2016). "'Fifty Shades' of Black and Red or How Carboxyl Groups Fine Tune Eumelanin and Pheomelanin Properties". International Journal of Molecular Sciences. 17 (5): 746. doi:10.3390/ijms17050746. PMC 4881568. PMID 27196900.
- Marklund, L; Moller MJ; Sandberg K; Andersson L (Dec 1996). "A missense mutation in the gene for melanocyte-stimulating hormone receptor (MC1R) is associated with the chestnut coat color in horses". Mammalian Genome. 7 (12): 895–9. doi:10.1007/s003359900264. PMID 8995760. S2CID 29095360.
- Wolf Horrell, Erin M.; Boulanger, Mary C.; D'Orazio, John A. (31 May 2016). "Melanocortin 1 Receptor: Structure, Function, and Regulation". Frontiers in Genetics. 7: 95. doi:10.3389/fgene.2016.00095. PMC 4885833. PMID 27303435.
- Rieder S, Taourit S, Mariat D, Langlois B, Guérin G (2001). "Mutations in the agouti (ASIP), the extension (MC1R), and the brown (TYRP1) loci and their association to coat color phenotypes in horses (Equus caballus)". Mammalian Genome. 12 (6): 450–455. doi:10.1007/s003350020017. PMID 11353392. S2CID 2012676.
- Voß, Katharina; Tetens, Julia; Thaller, Georg; Becker, Doreen (2020). "Coat Color Roan Shows Association with KIT Variants and No Evidence of Lethality in Icelandic Horses". Genes. 11 (6): 680. doi:10.3390/genes11060680. PMC 7348759. PMID 32580410.
- Fontanesi, L.; Tazzoli, M.; Beretti, F.; Russo, V. (October 2006). "Mutations in the melanocortin 1 receptor (MC1R) gene are associated with coat colours in the domestic rabbit (Oryctolagus cuniculus)". Animal Genetics. 37 (5): 489–493. doi:10.1111/j.1365-2052.2006.01494.x. PMID 16978179.
- Fontanesi, Luca; Beretti, Francesca; Riggio, Valentina; Dall'Olio, Stefania; González, Elena Gómez; Finocchiaro, Raffaella; Davoli, Roberta; Russo, Vincenzo; Portolano, Baldassare (December 2009). "Missense and nonsense mutations in melanocortin 1 receptor (MC1R) gene of different goat breeds: association with red and black coat colour phenotypes but with unexpected evidences". BMC Genetics. 10 (1): 47. doi:10.1186/1471-2156-10-47. PMC 2748843. PMID 19706191.
- Robbins, L.S.; Nadeau, J. H.; Johnson, K. R.; Kelly, M. A.; Roselli-Rehfuss, L.; Baack, E.; Mountjoy, K. G.; Cone, R. D. (1993). "Pigmentation phenotypes of variant extension locus alleles result from point mutations that alter MSH receptor function". Cell. 72 (6): 827–834. doi:10.1016/0092-8674(93)90572-8. PMID 8458079. S2CID 12179800.
- Joerg, H; Fries, H. R.; Meijerink, E.; Stranzinger, G. F. (1996). "Red coat color in Holstein cattle is associated with a deletion in the MSHR gene". Mammalian Genome. 7 (4): 317–318. doi:10.1007/s003359900090. PMID 8661706. S2CID 2497765.
- Newton, JM; Wilkie, A. L.; He, L.; Jordan, S. A.; Metallinos, D. L.; Holmes, N. G.; Jackson, I. J.; Barsh, G. S. (2000). "Melanocortin 1 receptor variation in the domestic dog". Mammalian Genome. 11 (1): 24–30. doi:10.1007/s003350010005. PMID 10602988. S2CID 1755908.
- Adalsteinsson, Stefán (January 1974). "Inheritance of the Palomino Color in Icelandic Horses". Journal of Heredity. 65 (1): 15–20. doi:10.1093/oxfordjournals.jhered.a108448. PMID 4847742.
- Wagner; Reissmann (2000). "New polymorphism detected in the horse MC1R gene". Animal Genetics. 31 (4): 289–290. doi:10.1046/j.1365-2052.2000.00655.x. PMID 11086549.
- Rendo F, Iriondo M, Manzano C, Estonba A (16 December 2009). "Identification of horse chestnut coat color genotype using SNaPshot". BMC Research Notes. 2: 255. doi:10.1186/1756-0500-2-255. PMC 2805676. PMID 20015355.
- Drogemuller, C; Giese, A.; Martins-Wess, F.; Wiedemann, S.; Andersson, L.; Brenig, B.; Fries, R.; Leeb, T (2006). "The mutation causing the black-and-tan pigmentation phenotype of Mangalitza pigs maps to the porcine ASIP locus but does not affect its coding sequence" (PDF). Mammalian Genome. 17 (1): 58–66. doi:10.1007/s00335-005-0104-1. PMID 16416091. S2CID 22542101.
- Locke, M. M.; Penedo, M. C. T.; Bricker, S. J.; Millon, L. V.; Murray, J. D. (October 2002). "Linkage of the grey coat colour locus to microsatellites on horse chromosome 25: Linkage mapping of horse grey locus". Animal Genetics. 33 (5): 329–337. doi:10.1046/j.1365-2052.2002.00885.x. PMID 12354140.
- Ludwig A, Pruvost M, Reissmann M, Benecke N, Brockmann GA, Castaños P, Cieslak M, Lippold S, Llorente L, Malaspinas AS, Slatkin M, Hofreiter M (2009-04-24). "Coat Color Variation at the Beginning of Horse Domestication". Science. 324 (5926): 485. Bibcode:2009Sci...324..485L. doi:10.1126/science.1172750. PMC 5102060. PMID 19390039.
- Mariat, Denis; Sead TAOURIT; Gérard GUÉRIN (2003). "A mutation in the MATP gene causes the cream coat colour in the horse". Genetics Selection Evolution. 35 (1). INRA, EDP Sciences: 119–133. doi:10.1051/gse:2002039. PMC 2732686. PMID 12605854.
- Sweet HO, Brilliant MH, Cook SA, Johnson KR, Davisson MT (1998). "A new allelic series for the underwhite gene on mouse chromosome 15". Journal of Heredity. 89 (6): 546–51. doi:10.1093/jhered/89.6.546. PMID 9864865.
- Wriedt-Ski, Chr. (December 1925). "Vererbungsuntersuchungen beim Pferd" [Heredity Studies in the Horse]. Zeitschrift für Induktive Abstammungs- und Vererbungslehre (in German). 37 (1): 88–101. doi:10.1007/BF01763328. S2CID 21565876.
- Cook, D; Brooks S; Bellone R; Bailey E (2008). Barsh, Gregory S. (ed.). "Missense Mutation in Exon 2 of SLC36A1 Responsible for Champagne Dilution in Horses". PLOS Genetics. 4 (9): e1000195. doi:10.1371/journal.pgen.1000195. PMC 2535566. PMID 18802473.
- Brooks; Lear; Adelson; Bailey (2007). "A chromosome inversion near the KIT gene and the Tobiano spotting pattern in horses". Cytogenetic and Genome Research. 119 (3–4): 225–230. doi:10.1159/000112065. PMID 18253033. S2CID 22835035.
- Mau, C.; Poncet, P. A.; Bucher, B.; Stranzinger, G.; Rieder, S. (2004). "Genetic mapping of dominant white (W), a homozygous lethal condition in the horse (Equus caballus)". Journal of Animal Breeding and Genetics. 121 (6): 374–383. doi:10.1111/j.1439-0388.2004.00481.x.
- Marklund, S.; Moller, M.; Sandberg, K.; Andersson, L. (1 March 1999). "Close association between sequence polymorphism in the KIT gene and the roan coat color in horses". Mammalian Genome. 10 (3): 283–288. doi:10.1007/s003359900987. PMID 10051325. S2CID 32790547.
- Hauswirth; Haase; Blatter (2012). "Mutations in MITF and PAX3 Cause "Splashed White" and Other White Spotting Phenotypes in Horses". PLOS Genetics. 8 (4): e1002653. doi:10.1371/journal.pgen.1002653. PMC 3325211. PMID 22511888.

harvard.edu

ui.adsabs.harvard.edu

Ludwig, Arne; Pruvost, Melanie; Reissmann, Monika; Benecke, Norbert; Brockmann, Gudrun A.; Castaños, Pedro; Cieslak, Michael; Lippold, Sebastian; Llorente, Laura; Malaspinas, Anna-Sapfo; Stalkin, Montgomery; Hofreiter, Michael (2009). "Coat Color Variation at the Beginning of Horse Domestication". Science. 324 (5926): 485. Bibcode:2009Sci...324..485L. doi:10.1126/science.1172750. PMC 5102060. PMID 19390039.

Wutke, Saskia; Benecke, Norbert; Sandoval-Castellanos, Edson; Döhle, Hans-Jürgen; Friederich, Susanne; Gonzalez, Javier; Hallsson, Jón Hallsteinn; Hofreiter, Michael; Lõugas, Lembi; Magnell, Ola; Morales-Muniz, Arturo; Orlando, Ludovic; Pálsdóttir, Albína Hulda; Reissmann, Monika; Ruttkay, Matej; Trinks, Alexandra; Ludwig, Arne (23 December 2016). "Spotted phenotypes in horses lost attractiveness in the Middle Ages". Scientific Reports. 6 (1): 38548. Bibcode:2016NatSR...638548W. doi:10.1038/srep38548. PMC 5141471. PMID 27924839.

Ludwig A, Pruvost M, Reissmann M, Benecke N, Brockmann GA, Castaños P, Cieslak M, Lippold S, Llorente L, Malaspinas AS, Slatkin M, Hofreiter M (2009-04-24). "Coat Color Variation at the Beginning of Horse Domestication". Science. 324 (5926): 485. Bibcode:2009Sci...324..485L. doi:10.1126/science.1172750. PMC 5102060. PMID 19390039.

nih.gov

pubmed.ncbi.nlm.nih.gov

"A horse of a different color: Genetics of camouflage and the dun pattern". ScienceDaily (Press release). December 21, 2015.
Ludwig, Arne; Pruvost, Melanie; Reissmann, Monika; Benecke, Norbert; Brockmann, Gudrun A.; Castaños, Pedro; Cieslak, Michael; Lippold, Sebastian; Llorente, Laura; Malaspinas, Anna-Sapfo; Stalkin, Montgomery; Hofreiter, Michael (2009). "Coat Color Variation at the Beginning of Horse Domestication". Science. 324 (5926): 485. Bibcode:2009Sci...324..485L. doi:10.1126/science.1172750. PMC 5102060. PMID 19390039.
Wutke, Saskia; Benecke, Norbert; Sandoval-Castellanos, Edson; Döhle, Hans-Jürgen; Friederich, Susanne; Gonzalez, Javier; Hallsson, Jón Hallsteinn; Hofreiter, Michael; Lõugas, Lembi; Magnell, Ola; Morales-Muniz, Arturo; Orlando, Ludovic; Pálsdóttir, Albína Hulda; Reissmann, Monika; Ruttkay, Matej; Trinks, Alexandra; Ludwig, Arne (23 December 2016). "Spotted phenotypes in horses lost attractiveness in the Middle Ages". Scientific Reports. 6 (1): 38548. Bibcode:2016NatSR...638548W. doi:10.1038/srep38548. PMC 5141471. PMID 27924839.
Rosengren Pielberg, Gerli; Golovko, Anna; Sundström, Elisabeth; Curik, Ino; Lennartsson, Johan; Seltenhammer, Monika H; Druml, Thomas; Binns, Matthew; Fitzsimmons, Carolyn; Lindgren, Gabriella; Sandberg, Kaj; Baumung, Roswitha; Vetterlein, Monika; Strömberg, Sara; Grabherr, Manfred; Wade, Claire; Lindblad-Toh, Kerstin; Pontén, Fredrik; Heldin, Carl-Henrik; Sölkner, Johann; Andersson, Leif (August 2008). "A cis-acting regulatory mutation causes premature hair graying and susceptibility to melanoma in the horse". Nature Genetics. 40 (8): 1004–1009. doi:10.1038/ng.185. PMID 18641652. S2CID 6666394.
- "Genetics Of White Horses Unraveled: One Mutation Makes Ordinary Horses Turn Grey, Then White, Very Young". ScienceDaily (Press release). July 23, 2008.
- Micillo, Raffaella; Panzella, Lucia; Koike, Kenzo; Monfrecola, Giuseppe; Napolitano, Alessandra; d'Ischia, Marco (17 May 2016). "'Fifty Shades' of Black and Red or How Carboxyl Groups Fine Tune Eumelanin and Pheomelanin Properties". International Journal of Molecular Sciences. 17 (5): 746. doi:10.3390/ijms17050746. PMC 4881568. PMID 27196900.
- Marklund, L; Moller MJ; Sandberg K; Andersson L (Dec 1996). "A missense mutation in the gene for melanocyte-stimulating hormone receptor (MC1R) is associated with the chestnut coat color in horses". Mammalian Genome. 7 (12): 895–9. doi:10.1007/s003359900264. PMID 8995760. S2CID 29095360.
- Wolf Horrell, Erin M.; Boulanger, Mary C.; D'Orazio, John A. (31 May 2016). "Melanocortin 1 Receptor: Structure, Function, and Regulation". Frontiers in Genetics. 7: 95. doi:10.3389/fgene.2016.00095. PMC 4885833. PMID 27303435.
- Rieder S, Taourit S, Mariat D, Langlois B, Guérin G (2001). "Mutations in the agouti (ASIP), the extension (MC1R), and the brown (TYRP1) loci and their association to coat color phenotypes in horses (Equus caballus)". Mammalian Genome. 12 (6): 450–455. doi:10.1007/s003350020017. PMID 11353392. S2CID 2012676.
- Voß, Katharina; Tetens, Julia; Thaller, Georg; Becker, Doreen (2020). "Coat Color Roan Shows Association with KIT Variants and No Evidence of Lethality in Icelandic Horses". Genes. 11 (6): 680. doi:10.3390/genes11060680. PMC 7348759. PMID 32580410.
- Fontanesi, L.; Tazzoli, M.; Beretti, F.; Russo, V. (October 2006). "Mutations in the melanocortin 1 receptor (MC1R) gene are associated with coat colours in the domestic rabbit (Oryctolagus cuniculus)". Animal Genetics. 37 (5): 489–493. doi:10.1111/j.1365-2052.2006.01494.x. PMID 16978179.
- Fontanesi, Luca; Beretti, Francesca; Riggio, Valentina; Dall'Olio, Stefania; González, Elena Gómez; Finocchiaro, Raffaella; Davoli, Roberta; Russo, Vincenzo; Portolano, Baldassare (December 2009). "Missense and nonsense mutations in melanocortin 1 receptor (MC1R) gene of different goat breeds: association with red and black coat colour phenotypes but with unexpected evidences". BMC Genetics. 10 (1): 47. doi:10.1186/1471-2156-10-47. PMC 2748843. PMID 19706191.
- Robbins, L.S.; Nadeau, J. H.; Johnson, K. R.; Kelly, M. A.; Roselli-Rehfuss, L.; Baack, E.; Mountjoy, K. G.; Cone, R. D. (1993). "Pigmentation phenotypes of variant extension locus alleles result from point mutations that alter MSH receptor function". Cell. 72 (6): 827–834. doi:10.1016/0092-8674(93)90572-8. PMID 8458079. S2CID 12179800.
- Joerg, H; Fries, H. R.; Meijerink, E.; Stranzinger, G. F. (1996). "Red coat color in Holstein cattle is associated with a deletion in the MSHR gene". Mammalian Genome. 7 (4): 317–318. doi:10.1007/s003359900090. PMID 8661706. S2CID 2497765.
- Newton, JM; Wilkie, A. L.; He, L.; Jordan, S. A.; Metallinos, D. L.; Holmes, N. G.; Jackson, I. J.; Barsh, G. S. (2000). "Melanocortin 1 receptor variation in the domestic dog". Mammalian Genome. 11 (1): 24–30. doi:10.1007/s003350010005. PMID 10602988. S2CID 1755908.
- Adalsteinsson, Stefán (January 1974). "Inheritance of the Palomino Color in Icelandic Horses". Journal of Heredity. 65 (1): 15–20. doi:10.1093/oxfordjournals.jhered.a108448. PMID 4847742.
- Wagner; Reissmann (2000). "New polymorphism detected in the horse MC1R gene". Animal Genetics. 31 (4): 289–290. doi:10.1046/j.1365-2052.2000.00655.x. PMID 11086549.
- Rendo F, Iriondo M, Manzano C, Estonba A (16 December 2009). "Identification of horse chestnut coat color genotype using SNaPshot". BMC Research Notes. 2: 255. doi:10.1186/1756-0500-2-255. PMC 2805676. PMID 20015355.
- Drogemuller, C; Giese, A.; Martins-Wess, F.; Wiedemann, S.; Andersson, L.; Brenig, B.; Fries, R.; Leeb, T (2006). "The mutation causing the black-and-tan pigmentation phenotype of Mangalitza pigs maps to the porcine ASIP locus but does not affect its coding sequence" (PDF). Mammalian Genome. 17 (1): 58–66. doi:10.1007/s00335-005-0104-1. PMID 16416091. S2CID 22542101.
- Locke, M. M.; Penedo, M. C. T.; Bricker, S. J.; Millon, L. V.; Murray, J. D. (October 2002). "Linkage of the grey coat colour locus to microsatellites on horse chromosome 25: Linkage mapping of horse grey locus". Animal Genetics. 33 (5): 329–337. doi:10.1046/j.1365-2052.2002.00885.x. PMID 12354140.
- Ludwig A, Pruvost M, Reissmann M, Benecke N, Brockmann GA, Castaños P, Cieslak M, Lippold S, Llorente L, Malaspinas AS, Slatkin M, Hofreiter M (2009-04-24). "Coat Color Variation at the Beginning of Horse Domestication". Science. 324 (5926): 485. Bibcode:2009Sci...324..485L. doi:10.1126/science.1172750. PMC 5102060. PMID 19390039.
- Mariat, Denis; Sead TAOURIT; Gérard GUÉRIN (2003). "A mutation in the MATP gene causes the cream coat colour in the horse". Genetics Selection Evolution. 35 (1). INRA, EDP Sciences: 119–133. doi:10.1051/gse:2002039. PMC 2732686. PMID 12605854.
- Sweet HO, Brilliant MH, Cook SA, Johnson KR, Davisson MT (1998). "A new allelic series for the underwhite gene on mouse chromosome 15". Journal of Heredity. 89 (6): 546–51. doi:10.1093/jhered/89.6.546. PMID 9864865.
- Cook, D; Brooks S; Bellone R; Bailey E (2008). Barsh, Gregory S. (ed.). "Missense Mutation in Exon 2 of SLC36A1 Responsible for Champagne Dilution in Horses". PLOS Genetics. 4 (9): e1000195. doi:10.1371/journal.pgen.1000195. PMC 2535566. PMID 18802473.
- Brooks; Lear; Adelson; Bailey (2007). "A chromosome inversion near the KIT gene and the Tobiano spotting pattern in horses". Cytogenetic and Genome Research. 119 (3–4): 225–230. doi:10.1159/000112065. PMID 18253033. S2CID 22835035.
- Marklund, S.; Moller, M.; Sandberg, K.; Andersson, L. (1 March 1999). "Close association between sequence polymorphism in the KIT gene and the roan coat color in horses". Mammalian Genome. 10 (3): 283–288. doi:10.1007/s003359900987. PMID 10051325. S2CID 32790547.
- Hauswirth; Haase; Blatter (2012). "Mutations in MITF and PAX3 Cause "Splashed White" and Other White Spotting Phenotypes in Horses". PLOS Genetics. 8 (4): e1002653. doi:10.1371/journal.pgen.1002653. PMC 3325211. PMID 22511888.
ncbi.nlm.nih.gov
- Pruvost, Melanie; Bellone, Rebecca; Benecke, Norbert; Sandoval-Castellanos, Edson; Cieslak, Michael; Kuznetsova, Tatyana; Morales-Muñiz, Arturo; O'Connor, Terry; Reissmann, Monika; Hofreiter, Michael; Ludwig, Arne (15 November 2011). "Genotypes of predomestic horses match phenotypes painted in Paleolithic works of cave art". Proceedings of the National Academy of Sciences. 108 (46): 18626–18630. Bibcode:2011PNAS..10818626P. doi:10.1073/pnas.1108982108. PMC 3219153. PMID 22065780.
- Imsland, Freyja; McGowan, Kelly; Rubin, Carl-Johan; Henegar, Corneliu; Sundström, Elisabeth; Berglund, Jonas; Schwochow, Doreen; Gustafson, Ulla; Imsland, Páll; Lindblad-Toh, Kerstin; Lindgren, Gabriella; Mikko, Sofia; Millon, Lee; Wade, Claire; Schubert, Mikkel; Orlando, Ludovic; Penedo, Maria Cecilia T; Barsh, Gregory S; Andersson, Leif (February 2016). "Regulatory mutations in TBX3 disrupt asymmetric hair pigmentation that underlies Dun camouflage color in horses". Nature Genetics. 48 (2): 152–158. doi:10.1038/ng.3475. PMC 4731265. PMID 26691985.
  - "A horse of a different color: Genetics of camouflage and the dun pattern". ScienceDaily (Press release). December 21, 2015.
  - Ludwig, Arne; Pruvost, Melanie; Reissmann, Monika; Benecke, Norbert; Brockmann, Gudrun A.; Castaños, Pedro; Cieslak, Michael; Lippold, Sebastian; Llorente, Laura; Malaspinas, Anna-Sapfo; Stalkin, Montgomery; Hofreiter, Michael (2009). "Coat Color Variation at the Beginning of Horse Domestication". Science. 324 (5926): 485. Bibcode:2009Sci...324..485L. doi:10.1126/science.1172750. PMC 5102060. PMID 19390039.
  - Wutke, Saskia; Benecke, Norbert; Sandoval-Castellanos, Edson; Döhle, Hans-Jürgen; Friederich, Susanne; Gonzalez, Javier; Hallsson, Jón Hallsteinn; Hofreiter, Michael; Lõugas, Lembi; Magnell, Ola; Morales-Muniz, Arturo; Orlando, Ludovic; Pálsdóttir, Albína Hulda; Reissmann, Monika; Ruttkay, Matej; Trinks, Alexandra; Ludwig, Arne (23 December 2016). "Spotted phenotypes in horses lost attractiveness in the Middle Ages". Scientific Reports. 6 (1): 38548. Bibcode:2016NatSR...638548W. doi:10.1038/srep38548. PMC 5141471. PMID 27924839.
  - Micillo, Raffaella; Panzella, Lucia; Koike, Kenzo; Monfrecola, Giuseppe; Napolitano, Alessandra; d'Ischia, Marco (17 May 2016). "'Fifty Shades' of Black and Red or How Carboxyl Groups Fine Tune Eumelanin and Pheomelanin Properties". International Journal of Molecular Sciences. 17 (5): 746. doi:10.3390/ijms17050746. PMC 4881568. PMID 27196900.
  - Wolf Horrell, Erin M.; Boulanger, Mary C.; D'Orazio, John A. (31 May 2016). "Melanocortin 1 Receptor: Structure, Function, and Regulation". Frontiers in Genetics. 7: 95. doi:10.3389/fgene.2016.00095. PMC 4885833. PMID 27303435.
  - Voß, Katharina; Tetens, Julia; Thaller, Georg; Becker, Doreen (2020). "Coat Color Roan Shows Association with KIT Variants and No Evidence of Lethality in Icelandic Horses". Genes. 11 (6): 680. doi:10.3390/genes11060680. PMC 7348759. PMID 32580410.
  - Fontanesi, Luca; Beretti, Francesca; Riggio, Valentina; Dall'Olio, Stefania; González, Elena Gómez; Finocchiaro, Raffaella; Davoli, Roberta; Russo, Vincenzo; Portolano, Baldassare (December 2009). "Missense and nonsense mutations in melanocortin 1 receptor (MC1R) gene of different goat breeds: association with red and black coat colour phenotypes but with unexpected evidences". BMC Genetics. 10 (1): 47. doi:10.1186/1471-2156-10-47. PMC 2748843. PMID 19706191.
  - Rendo F, Iriondo M, Manzano C, Estonba A (16 December 2009). "Identification of horse chestnut coat color genotype using SNaPshot". BMC Research Notes. 2: 255. doi:10.1186/1756-0500-2-255. PMC 2805676. PMID 20015355.
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