Beardsley, P. M.; Olmstead, R. G. (2002). "Redefining Phrymaceae: the placement of Mimulus, tribe Mimuleae, and Phryma". American Journal of Botany. 89 (7): 1093–1102. doi:10.3732/ajb.89.7.1093. JSTOR4122195. PMID21665709.
Cooley, Arielle M.; Willis, John H. (2009). "Genetic divergence causes parallel evolution of flower color in Chilean Mimulus". New Phytologist. 183 (3): 729–739. doi:10.1111/j.1469-8137.2009.02858.x. PMID19453433.
Beardsley, P. M.; Olmstead, R. G. (2002). "Redefining Phrymaceae: the placement of Mimulus, tribe Mimuleae, and Phryma". American Journal of Botany. 89 (7): 1093–1102. doi:10.3732/ajb.89.7.1093. JSTOR4122195. PMID21665709.
Beardsley, P. M.; Olmstead, R. G. (2002). "Redefining Phrymaceae: the placement of Mimulus, tribe Mimuleae, and Phryma". American Journal of Botany. 89 (7): 1093–1102. doi:10.3732/ajb.89.7.1093. JSTOR4122195. PMID21665709.
Cooley, Arielle M.; Willis, John H. (2009). "Genetic divergence causes parallel evolution of flower color in Chilean Mimulus". New Phytologist. 183 (3): 729–739. doi:10.1111/j.1469-8137.2009.02858.x. PMID19453433.