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Kodandaramaiah, U. (2011). "Eyespot evolution: phylogenetic insights from Junonia and related butterfly genera (Nymphalidae: Junoniini)". Behavioral Ecology. 22 (5): 1264–1271. doi:10.1111/j.1525-142X.2009.00357.x. PMID19754706. S2CID1832497.
Breakfield, M.P.; Gates, J.; Keys, D.; Kesbeke, F.; Wijngaarden, J. P.; Monteiro, A.; French, V.; Carroll, S. B. (November 1996). "Development, plasticity and evolution of butterfly eyespot patterns". Nature. 384 (6606): 236–242. Bibcode:1996Natur.384..236B. doi:10.1038/384236a0. PMID12809139. S2CID3341270.
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Kodandaramaiah, U. (2011). "Eyespot evolution: phylogenetic insights from Junonia and related butterfly genera (Nymphalidae: Junoniini)". Behavioral Ecology. 22 (5): 1264–1271. doi:10.1111/j.1525-142X.2009.00357.x. PMID19754706. S2CID1832497.
Sourakov, Andrei (2013). "Two heads are better than one: false head allows Calycopis cecrops (Lycaenidae) to escape predation by a Jumping Spider, Phidippus pulcherrimus (Salticidae)". Journal of Natural History. 47 (15–16): 1047–1054. Bibcode:2013JNatH..47.1047S. doi:10.1080/00222933.2012.759288. S2CID84454608.
Robbins, Robert K. (November 1981). "The 'False Head' Hypothesis: Predation and Wing Pattern Variation of Lycaenid Butterflies". The American Naturalist. 118 (5): 770–775. doi:10.1086/283868. S2CID34146954.
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