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Nickol, B. B.; Heard, R. W.; Smith, N. F. (2002). "Acanthocephalans from crabs in the southeastern US, with the first intermediate hosts known for Arhythmorhynchus frassoni and Hexaglandula corynosoma". Journal of Parasitology. 88 (1): 79–83. doi:10.1645/0022-3395(2002)088[0079:afcits]2.0.co;2. PMID12053984.