Oda, Y. 2002. Photosynthetic characteristics and geographical distribution of diploid Fragaria species native in Japan. Acta Hort. 567: 381–384. AbstractArchived 2018-06-02 at the Wayback Machine.
Hummer, K. E., Sabitov, A., & Davis, T. 2005. Iturup and Sakhalin Island strawberries. HortScience 40(4): 1127. Abstract.
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Oda, Y. 2002. Photosynthetic characteristics and geographical distribution of diploid Fragaria species native in Japan. Acta Hort. 567: 381–384. AbstractArchived 2018-06-02 at the Wayback Machine.