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Hentschel, Jörn; von Konrat, Matthew J.; Pócs, Tamás; Schäfer-Verwimp, Alfons; Jonathan Shaw, A.; Schneider, Harald; Heinrichs, Jochen (1 July 2009). "Molecular insights into the phylogeny and subgeneric classification of Frullania Raddi (Frullaniaceae, Porellales)". Molecular Phylogenetics and Evolution. 52 (1): 142–156. doi:10.1016/j.ympev.2008.12.021. ISSN1055-7903.