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Samigullin, T.K.; Martin, W.F.; Troitsky, A.V.; Antonov, A.S. (1999). "Molecular data from the chloroplast rpoC1 gene suggest a deep and distinct dichotomy of contemporary spermatophytes into two monophyla: gymnosperms (including Gnetalaes) and angiosperms". Journal of Molecular Evolution. 49 (3): 310–315. Bibcode:1999JMolE..49..310S. doi:10.1007/PL00006553. PMID10473771. S2CID4232968.
Rydin, C.; Kallersjo, M.; Friist, E.M. (2002). "Seed plant relationships and the systematic position of Gnetales based on nuclear and chloroplast DNA: Conflicting data, rooting problems, and the monophyly of conifers". International Journal of Plant Sciences. 163 (2): 197–214. doi:10.1086/338321. JSTOR3080238. S2CID84578578.
Gugerli, F.; Sperisen, C.; Buchler, U.; Brunner, L.; Brodbeck, S.; Palmer, J.D.; Qiu, Y.L. (2001). "The evolutionary split of Pinaceae from other conifers: evidence from an intron loss and a multigene phylogeny". Molecular Phylogenetics and Evolution. 21 (2): 167–175. doi:10.1006/mpev.2001.1004. PMID11697913.
Friis, Else Marie; Crane, Peter R.; Pedersen, Kaj Raunsgaard; Bengtson, Stefan; Donoghue, Philip C.J.; Grimm, Guido W.; Stampanoni, Marco (November 2007). "Phase-contrast X-ray microtomography links Cretaceous seeds with Gnetales and Bennettitales". Nature. 450 (7169): 549–552. Bibcode:2007Natur.450..549F. doi:10.1038/nature06278. ISSN0028-0836. PMID18033296. S2CID1198220.
Hajibabaei, M.; Xia, J.; Drouin, G. (2006). "Seed plant phylogeny: Gnetophytes are derived conifers and a sister group to Pinaceae". Molecular Phylogenetics and Evolution. 40 (1): 208–217. doi:10.1016/j.ympev.2006.03.006. PMID16621615.
Samigullin, T.K.; Martin, W.F.; Troitsky, A.V.; Antonov, A.S. (1999). "Molecular data from the chloroplast rpoC1 gene suggest a deep and distinct dichotomy of contemporary spermatophytes into two monophyla: gymnosperms (including Gnetalaes) and angiosperms". Journal of Molecular Evolution. 49 (3): 310–315. Bibcode:1999JMolE..49..310S. doi:10.1007/PL00006553. PMID10473771. S2CID4232968.
Friis, Else Marie; Crane, Peter R.; Pedersen, Kaj Raunsgaard; Bengtson, Stefan; Donoghue, Philip C.J.; Grimm, Guido W.; Stampanoni, Marco (November 2007). "Phase-contrast X-ray microtomography links Cretaceous seeds with Gnetales and Bennettitales". Nature. 450 (7169): 549–552. Bibcode:2007Natur.450..549F. doi:10.1038/nature06278. ISSN0028-0836. PMID18033296. S2CID1198220.
Qiu, Y.L.; Lee, J.; Bernasconi-Quadroni, F.; Soltis, D.E.; Soltis, P.S.; Zanis, M.; Zimmer, E.A.; Chen, Z.; Savalainen, V. & Chase, M.W. (1999). "The earliest angiosperms: evidence from mitochondrial, plastid and nuclear genomes". Nature. 402 (6760): 404–407. Bibcode:1999Natur.402..404Q. doi:10.1038/46536. PMID10586879. S2CID4380796.
Rydin, C.; Kallersjo, M.; Friist, E.M. (2002). "Seed plant relationships and the systematic position of Gnetales based on nuclear and chloroplast DNA: Conflicting data, rooting problems, and the monophyly of conifers". International Journal of Plant Sciences. 163 (2): 197–214. doi:10.1086/338321. JSTOR3080238. S2CID84578578.
Coiro, M.; Chomicki, G.; Doyle, J.A. (2018). "Experimental signal dissection and method sensitivity analyses reaffirm the potential of fossils and morphology in the resolution of the relationship of angiosperms and Gnetales". Paleobiology. 44 (3): 490–510. Bibcode:2018Pbio...44..490C. doi:10.1017/pab.2018.23. S2CID91488394.
Samigullin, T.K.; Martin, W.F.; Troitsky, A.V.; Antonov, A.S. (1999). "Molecular data from the chloroplast rpoC1 gene suggest a deep and distinct dichotomy of contemporary spermatophytes into two monophyla: gymnosperms (including Gnetalaes) and angiosperms". Journal of Molecular Evolution. 49 (3): 310–315. Bibcode:1999JMolE..49..310S. doi:10.1007/PL00006553. PMID10473771. S2CID4232968.
Stull, Gregory W.; Qu, Xiao-Jian; Parins-Fukuchi, Caroline; Yang, Ying-Ying; Yang, Jun-Bo; Yang, Zhi-Yun; Hu, Yi; Ma, Hong; Soltis, Pamela S.; Soltis, Douglas E.; Li, De-Zhu; Smith, Stephen A.; Yi, Ting-Shuang (2021). "Gene duplications and phylogenomic conflict underlie major pulses of phenotypic evolution in gymnosperms". Nature Plants. 7 (8): 1015–1025. doi:10.1038/s41477-021-00964-4. PMID34282286. S2CID236141481.