Turner, J R G (2013). "The dawn flight of the gold swift Hepialus hecta: predator avoidance and the integration of complex lek behaviour (Lepidoptera, Hepialidae)". Biological Journal of the Linnean Society. 110 (2): 305–319. doi:10.1111/bij.12145.
Turner, J R G (2015). "The flexible lek: Phymatopus hecta the gold swift demonstrates the evolution of leking and male swarming via a hotspot (Lepidoptera: Hepialidae)". Biological Journal of the Linnean Society. 114: 184–201. doi:10.1111/bij.12411.
Mallet, J (1984). "Sex roles in the ghost moth Hepialus humuli (L.) and a review of mating in the Hepialidae (Lepidoptera)". Zoological Journal of the Linnean Society. 79: 67-82. doi:10.1111/j.1096-3642.1984.tb02320.x.