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McRae, W. Alan; Landers, J. Larry; Garner, James A. (July 1981). "Movement Patterns and Home Range of the Gopher Tortoise". American Midland Naturalist. 106 (1): 165. doi:10.2307/2425146. JSTOR2425146.
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Palmer, B. D.; Guillette, L. J. (November 1988). "Histology and functional morphology of the female reproductive tract of the tortoise Gopherus polyphemus". American Journal of Anatomy. 183 (3): 200–211. doi:10.1002/aja.1001830303. ISSN0002-9106. PMID3213826.
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Keddy, P.A. (2009). "Thinking big: A conservation vision for the Southeastern coastal plain of North America". Southeastern Naturalist. 8 (2): 213–226. doi:10.1656/058.008.0202. S2CID73678050.
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McGuire, Jessica L.; Smith, Lora L.; Guyer, Craig; Yabsley, Michael J. (1 October 2014). "Effects of Mycoplasmal Upper-Respiratory-Tract Disease on Movement and Thermoregulatory Behavior of Gopher Tortoises (Gopherus Polyphemus) in Georgia, USA". Journal of Wildlife Diseases. 50 (4): 745–756. doi:10.7589/2013-11-306. ISSN0090-3558. PMID25121399. S2CID5470163.
Palmer, B. D.; Guillette, L. J. (November 1988). "Histology and functional morphology of the female reproductive tract of the tortoise Gopherus polyphemus". American Journal of Anatomy. 183 (3): 200–211. doi:10.1002/aja.1001830303. ISSN0002-9106. PMID3213826.
Quinn, Daniel P.; Buhlmann, Kurt A.; Jensen, John B.; Norton, Terry M.; Tuberville, Tracey D. (September 2018). "Post-release movement and survivorship of head-started gopher tortoises". The Journal of Wildlife Management. 82 (7): 1545–1554. Bibcode:2018JWMan..82.1545Q. doi:10.1002/jwmg.21493. ISSN0022-541X.
Diemer, Joan E. (1986). "The Ecology and Management of the Gopher Tortoise in the Southeastern United States". Herpetologica. 42 (1): 125–133. ISSN0018-0831. JSTOR3892243.